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Paleobiota of the Ciechocinek Formation

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Reconstructed picture of dinosaurs
Terrestrial environment of the Toarcian Fennoscandinavia, with flora based on the Sorthat Formation. Dinosaurs are based on material found on various locations of the German realm of the Ciechocinek Formation

The Ciechocinek Formation is a Jurassic (lower to middle Toarcian) geologic formation which extends across the Baltic coast from Grimmen, Germany, to Nida, Lithuania, with its major sequence in Poland and boreholes in Kaliningrad. Dinosaur species uncovered here, including Emausaurus and other unclassified genus.

In Poland, the main basin lacks marine microfauna. The Ciechocinek Formation in the Częstochowa-Zawiercie area reveals the remains of a wide range of prehistoric environments; the Fore-Sudetic Monocline region must have been an extensive bay similar to Lake Maracaibo in Venezuela. The basin's shore zone was a flat, muddy, marshy coastal plain.

The region has the remains of the Wrêczyca River, which was active for most of the Pliensbachian/Toarcian period. At the Brody-Lubienia borehole (Lubienia), which once formed part of the river's east side, an alluvial system ended at a delta and discharged into a shallow marine bay and lagoon. A number of phyllopods and fossilized plant roots have been found here, where they were discharged by the river. Paleosol indicates that the lagoon had a maximum depth of about 6 metres (20 ft).

Foraminifera

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Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Genus Species Location Material Abundance Notes Images

Ammobaculites[1]

  • A. vetusta
  • A. linea
  • Mechowo 1 Borehole

Shells

Abundant

A marine foraminiferan, member of the family Ammomarginulininae inside Lituolidae

Ammodiscus[2][3][4][5]

  • A. glumaceous
  • A. orbis
  • A. cf. orbis
  • Pabianice
  • Łutowiec
  • Żarki
  • Wyżyna Krakowsko-Częstochowska

Shells

Rare

A marine foraminiferan, type member of the family Ammodiscinae inside Ammodiscina. Benthic foraminiferal successions heavily dominated by Ammodiscus and Trochammina are known from several Late Triassic to Middle Jurassic sediment packages along the Atlantic margin of northwestern Europe.

Berthelinella[6]

  • B. paradoxa
  • Dobbertin

Shells

Very scarce

A marine/brackish foraminiferan, member of the family Plectofrondiculariidae inside Polymorphinina

Bolivina[6]

  • B. liasica
  • Dobbertin

Shells

Abundant

A marine/brackish foraminiferan, type member of the family Bolivinidae inside Bolivinoidea; reaches 40% of sampled foranimiferans on at least one sample

Citharina[4][5]

  • C. sp.
  • Boża Wola
  • Gorzów Wiepolski

Shells

Rare

A marine foraminiferan, type member of the family Vaginulininae inside Nodosarioidea

Cornuspira[6]

  • C. orbicula
  • Dobbertin

Shells

Present in low numbers

A marine/brackish foraminiferan, type member of the family Cornuspirinae inside Cornuspiroidea

Crithionina[1]

  • C. sp.
  • Aleksandrów I Borehole
  • Gorzów Wiepolski
  • Mechowo 1 Borehole

Shells

Rare

A marine foraminiferan, type member of the family Saccamminidae inside Astrorhizacea

Dentalina[6]

  • D. deslongchampsi
  • D. primaeva
  • D. pseudocommunis
  • D. submucronata
  • D. subtenuicollis
  • D. tenuistriata
  • D. torta
  • D. vetusta
  • Pabianice
  • Łutowiec
  • Żarki
  • Dobbertin

Shells

Diverse but scarce

A marine/brackish foraminiferan, member of the family Nodosariidae inside Nodosariinae

Eoguttulina[6]

  • E. bilocularis
  • E. liassica
  • E. cf.simplex
  • Dobbertin

Shells

Scarce

A marine/brackish foraminiferan, member of the family Polymorphininae inside Polymorphinina

Falsopalmula[6]

  • F. insignis
  • Dobbertin

Shells

Very scarce

A marine/brackish foraminiferan, member of the family Robuloididae inside Robuloidoidea

Frondicularia[6]

  • F. bicostata
  • F. sulcata
  • F. terquemi
  • F. sp.
  • Dobbertin

Shells

Diverse but scarce

A marine/brackish foraminiferan, member of the family Nodosariidae inside Nodosariinae

Glomospira[6][2][3][4][5]

  • G. gordialis
  • G. sp.
  • Boża Wola
  • Wyżyna Krakowsko-Częstochowska
  • Dobbertin

Shells

Moderately present

A marine/brackish foraminiferan, member of the family Usbekistaniinae inside Ammodiscidae; its abundance is lower than other genera, yet reaches peaks of 5-10% in at least one sample

Haplophragmoides[1][4][5]

  • H. tryssa
  • H. platus
  • H. sp.
  • Aleksandrów I Borehole
  • Gorzów Wiepolski
  • Mechowo 1 Borehole
  • Wyżyna Krakowsko-Częstochowska

Shells

Diverse but rare

A marine foraminiferan, member of the family Lituoloidea inside Lituolida

Involutina[1]

  • I. silicea
  • Aleksandrów I Borehole
  • Krzyzskie

Shells

Rare

A marine foraminiferan, member of the family Ammodiscinae inside Ammodiscina

Lagena[6]

  • L. globosa
  • L. oxystoma
  • L. globosa
  • L. tenuicostata
  • L. sp.
  • Dobbertin

Shells

Scarce

A marine/brackish foraminiferan, member of the family Nodosariidae inside Nodosariinae

Lagenammina[2][3]

  • L. sp.
  • Pabianice
  • Łutowiec
  • Żarki

Shells

Rare

A marine foraminiferan, member of the family Saccamminidae inside Astrorhizacea; sometimes confused with the genus Arlagenammum

Lenticulina[6][1]

  • L. acutiangulata
  • L. müensteri
  • L. varians
  • L. sublaevis
  • Boża Wola
  • Wyżyna Krakowsko-Częstochowska
  • Dobbertin

Shells

Diverse but very scarce

A marine foraminiferan, member of the family Vaginulinidae inside Nodosarioidea

Lingulina[6]

  • L. tenera
  • Dobbertin

Shells

Abundant

A marine/brackish foraminiferan, type member of the family Lingulininae inside Nodosariinae; reaches percentages of 25% in some samples

Marginulina[6]

  • M. prima
  • M. simplex
  • M. oolithica
  • Dobbertin

Shells

Abundant

A marine/brackish foraminiferan, type member of the family Marginulininae inside Vaginulinidae; Marginulina prima reaches a 35% on a few samples

Nodosaria[6]

  • N. dispar
  • N. agglutinans
  • N. elongata
  • N. germanica
  • N. hirsuta
  • N. dispar
  • N. mitis
  • N. regularis
  • N. simplex
  • Dobbertin

Shells

Diverse but scarce

A marine/brackish foraminiferan, type member of the family Nodosariidae inside Nodosariinae. Despite its diversity, it is very scarce on most samples, with less than 1-2% of presence.

Ophtalmidium[6][4][5]

  • O. orbiculare
  • O. carinatum
  • O. sp.
  • Boża Wola
  • Dobbertin

Shells

Present in low numbers

A marine/brackish foraminiferan, type member of the family Ophthalmidiidae inside Cornuspiroidea

Placopsilina[1]

  • P. complanata
  • Mechowo 1 Borehole

Shells

Rare

A marine foraminiferan, member of the family Lituoloidea inside Lituolida

Rectoglandulina[6]

  • R. oviformis
  • R. sexcostata
  • Dobbertin

Shells

Scarce

A marine/brackish foraminiferan, member of the family Nodosariidae inside Nodosariinae

Saccammina[2][3]

  • S.? sp
  • Pabianice
  • Łutowiec
  • Żarki

Shells

Rare

A marine foraminiferan, type member of the family Saccamminidae inside Astrorhizacea. Sometimes confused with the genus Arsaccammum or called Saccamina. The local specimens are rather rare, incomplete, and complicated to identify.

Stilostomella[6]

  • S.? sp.
  • Dobbertin

Shells

One specimen

A marine/brackish foraminiferan, type member of the family Stilostomellidae inside Nodosariinae

Trochamminoides[6]

  • T. sp.
  • Dobbertin

Shells

Very scarce

A marine/brackish foraminiferan, member of the family Trochamminoidae inside Lituolida

Trochammina[2][3][4][5]

  • T. sp
  • T.? sp
  • Pabianice
  • Łutowiec
  • Boża Wola
  • Żarki
  • Wyżyna Krakowsko-Częstochowska

Shells

Rare

A marine foraminiferan, type member of the family Trochammininae inside Trochamminina. Small-sized AmmodiscusTrochammina assemblages are found associated with delta-influenced shelf environment, where biota would have been stressed by intermittent periods with moderate hypoxia combined with lowered salinity and storm impacts.

Turrispirillina[6]

  • T. conoidea
  • Dobbertin

Shells

Very scarce

A marine/brackish foraminiferan, member of the family Spirillinidae inside Spirillinina

Vaginulina[6]

  • V. anomala
  • Dobbertin

Shells

Scarce

A marine/brackish foraminiferan, type member of the family Vaginulininae inside Vaginulinidae

Dinoflagellates

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Genus Species Location Material Abundance Notes Images

Luehndea[7]

  • L. spinosa
  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Dinocysts

Abundant

A marine Dinophyceae Dinoflagellatan, type member of the family Luehndeoideae inside Mancodiniaceae. Presence of Luehndea spinosa suggests Late Pliensbachian–earliest Toarcian age of studied assemblages. The marine dinoflagellate cyst Luehndea spinosa and foraminiferal linings were found only in the lower half of the Kozłowice succession.

Nannoceratopsis[7]

  • N. senex
  • N. triceras
  • N. raunsgardii
  • N. ridingii
  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Dinocysts

Dominant

A marine Dinophyceae Dinoflagellatan, type member of the family Nannoceratopsiaceae inside Nannoceratopsiales. The large amount of Cysts of the genus point to more diversified marine palaeoenvironments. N. senex is the most abundant. Locally Nannoceratopsis recovers series of marine transgressions and regressions, pointing to the presence of interbedding marine and brackish sediments.

Fungi

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Genus Species Location Material Notes Images

Xylophagous Fungi[8]

  • Morphotype A (Aseptae)
  • Morphotype B (Sphaerical)
  • Morphotype C (Sack-shaped)
  • Morphotype D (Transverse septa)
  • Morphotype E (Amerospore)
  • Morphotype F (Phragmospores)
  • Morphotype G (Dictyospores)
  • Brody-Lubienia borehole.[8]
  • Gorzów Wielkopolski Borehole
  • Mechowo Borehole
  • Parkoszowice borehole
  • Fungal Spores
  • Hyphae-like attachments

Saprophyte fungal Spores from Marine and Deltaic Settings associated with wood and Litter.[8] The frequency of fungal spores on the Polish basin on the Pliensbachian-Toarcian transition is correlated with negative C isotope peaks and enhanced cuticular plant litter accumulation, pointing to the climate-driven enhanced decomposition of wood and rapid destruction of terrestrial carbon, which may have played an important role in the aggravation of the Jurassic Greenhouse disaster.[8] Associated with a high rate of organic burial, the presence of Fungal Matter increased on the Uppermost layers of the Drzewica Formation, with a continued deposition between the T-OAE extinction, and several ups and downs on the Ciechocinek Formation, related with local climate and humidity changes. This is rather a reflection of the efficiency of terrestrial biodegradation.[8] Measured increasing of temperature favored local fungal-mediated decomposition of plant litter, specifically of normally resistant wood.[8] Observed fungal spores represent various morphotypes and resemble superficially other palynogenic detritus, such as spherical Prasinophyceae.[8] In some cases spherical fungal spores show structures related to Hyphae attachments.[8]

Spores found on the Polish are mostly from Saprophyte (wood decomposers) fungi, resembling the extant genera like Scutellinia, which are probably the main origin for the Fungal Spores found on the Ciechocinek Formation

Invertebrates

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Ichnofossils

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Genus Species Type Location Material Origin Notes Images

Chondrites[9][10]

  • C. bollensis

Fodinichnia

  • Dobbertin Clay Pit

Small branching burrows

Saltwater/Blackish burrow-like ichnofossils. Interpreted as the feeding burrow of a sediment-ingesting animal. A more recent study has found that Scoloplos armiger and Heteromastus filiformis, occurring in the German Wadden Sea in the lower parts of tidal flats, make burrows that are homonymous with numerous trace fossils of the ichnogenus.

Illustration of Chondrites bollensis

Diplocraterion[11]

  • D. parallelum

Domichnia

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard
  • Parkoszowice 58 BN borehole
  • Żuki ul. 88-2 Borehole
  • Gorzków BN Borehole
  • Bobolice 68-km Borehole
  • Choroń 31-BN Borehole

U-shaped burrows

Marine-Mangroove Vertical, U-shaped, single-spreite Burrows; unidirectional or bidirectional spreite, generally continuous, rarely discontinuous. Most Diplocraterion show only protrusive spreit, like the local ones, produced under predominantly erosive conditions where the organism was constantly burrowing deeper into the substrate as sediment was eroded from the top.

Diplocraterion parallelum diagram

Gyrochorte[11]

  • G. isp.

Fodinichnia

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Winding, horizontal, double ridge burrows, separated by a median groove.

Saltwater/Blackish burrow-like ichnofossils. On the Kozlowice strata, only a few specimens were observed. Gyrochorte is interpreted as a result of active digging on the sediment by a deposit-feeding worm-like animal, probably Annelid or similar kinds of creatures.[11]

Helminthopsis[11]

  • H. isp.

Fodinichnia

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Simple, unbranched, horizontal cylinder traces

Saltwater/Blackish burrow-like ichnofossils. There are a few specimens in the Kozlowice outcrop. It is interpreted as a grazing trail or Fodinichnia, produced at shallow depth in sediment by Polychaetes and Priapulids.[11]

Example of Helminthopsis fossil

Megagrapton[10]

  • M. isp.

Pascichnia

  • Dobbertin Clay Pit

Cylindrical strands with branches

  • Polychaetes
  • Sipunculans
  • Enteropneustans
  • Echiurans

Saltwater/Blackish trace ichnofossils. Probably done by Polychaetes

Palaeophycus[11]

  • P. tubularis

Domichnia

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Straight or gently curved tubular burrows.

  • Polychaetes
  • Semiaquatic Insects (Orthoptera and Hemiptera)
  • Semiaquatic and non-aquatic Beetles.

Saltwater/Blackish burrow-like ichnofossils. Palaeophycus is less common than Planolites in deposits of the Ciechocinek Formation. On the Kozlowice outcrop however there are numerous specimens occur, interpreted as the result of passive filling of polychaete burrows.[11]

Example of Palaeophycus fossil

Planolites[11]

  • P. montanus
  • P. beverleyensis

Pascichnia

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Cylindrical or elliptical curved/tortuous trace fossils

  • Polychaetes
  • Insects

Saltwater/Blackish burrow-like ichnofossils. Planolites is really common in all types of the Ciechocinek Formation deposits. It is referred to vermiform deposit-feeders, mainly Polychaetes, producing active Fodinichnia. It is controversial, since is considered a strictly a junior synonym of Palaeophycus.

Example of Planolites fossil

Protovirgularia[11]

  • P. isp.

Pasichnia

  • Kozłowice Clay Pit
  • "Boroszów" Brickyard

Bilobate trace fossil

  • Branchiopods
  • Bivalves

Saltwater/Blackish and Freshwater bottom Trace Fossils. Protovirgularia is a Repichnia form, ascribed to the activity of Bivalves, leaving a trace due to the rhythmic action of a foot.[11]

Skolithos[12]

  • S. isp.

Domichnia

  • Müssentin gravel pit

Cylindrical strands with branches

  • Polychaetes
  • Phoronidans

Blackish trace ichnofossils. Interpreted as dwelling structures of vermiform animals, more concretely the Domichnion of a suspension-feeding Worm or Phoronidan.

Spongeliomorpha[11][13]

  • S. isp.
  • Repichnia
  • Fodichnia
  • Kozłowice Clay Pit
  • "Boroszów" Brickyard
  • Pawłowice 40 Borehole

Horizontal and subhorizontal, branching tunnels, cylindrical or elliptical in cross-section, displaying elongated striation on exterior of burrow casts

Saltwater/Blackish burrow-like ichnofossils. On the Kozłowice section a series of sand-filled tunnels of Spongeliomorpha are preserved as Endichnia within mudstone, preserved as Tubular Tempestites (Storm-burrow filling), open tubes produced by animals burrowing in a stable, stiff or firm substrate.[13] Here the tubes occur in one horizon with thin sandy laminae and lenses that represent isolated Starved Ripples carried on the muddy sea-floor by storm-generated Traction power network.[13]

Taenidium[9][10]

  • T. serpentinum

Fodinichnia

  • Dobbertin Clay Pit

Unlined meniscate burrows

Saltwater/Blackish burrow-like ichnofossils. Taenidium is a meniscate backfill structure, usually considered to be produced by an animal progressing axially through the sediment and depositing alternating packets of differently constituted sediment behind it as it moves forward.

Teichichnus[14]

  • Teichichnus isp.

Fodinichnia

  • Gorzów Wielkopolski IG 1 borehole

Vertical to oblique, unbranched or branched, elongated to arcuate spreite burrow

Saltwater/Blackish burrow-like ichnofossils. Is common on the Pomerania Region. The overall morphology and details of the burrows, in comparison with modern analogues and neoichnological experiments, suggest Echiurans (spoon worms) or Holothurians (sea cucumbers) with a combined suspension- and deposit-feeding behaviour as potential producers.

Teichichnus burrows

Annelida

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Genus Species Stratigraphic position Material Notes Images

Dictyothylakos[15]

  • D. pesslerae
  • D. "sp. Sing.1964"
  • Brody-Lubienia borehole
  • Wolin IG-1 borehole
  • Kaszewy 1 borehole
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole

Cocoons

Freshwater Clitellata Cocoons (Oligochaeta and Hirudinea), identified with palynological residues, and through to be tridimensional nets of probable algal origin.[16] Fragmentary mesh-like networks of happiness threads composed of homogenous translucent material. They show the outer wall (hapsine) construction specific to clitellate annelids and lack an alytine (inner) layer. A disorderly meshwork of the hapsine layer and hapsine fibers of unequal thickness, are diagnostic of the type species Dictyothylakos pesslerae. The cocoons Dictyothylakos pesslerae resemble specially those of modern Leechs, and are common on flooded basin sediments, which implies not only the presence of parasitic leeches, but also the presence of large hosts nearby, as has been confirmed on the case of the Ciechocinek Formation, thanks to the presence of not only dinosaurs but also Dipnoi and other freshwater taxa.

Example of leech cocoon
Placobdella, example of leech

Brachiopoda

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Genus Species Stratigraphic position Material Notes Images

Discina[17][18]

  • D. cornucopiae
  • D. papyracea
  • D. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

Cunchs

A Saltwater Brachiopodan, member of Discinidae inside Discinida. The classification of the Discinidae is rather treated with confusion, due to the description and the identification of either extinct and extant genera and species. The shells of this genus are the only identifiable brachiopods of the formation.

Bivalvia

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Genus Species Stratigraphic position Material Notes Images

Bositra[17][18][19]

  • B. buchii
  • B. radiata
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Ahrensburg

Shells

A Saltwater Oyster, member of "Posidoniidae" inside Ostreoida. A genus classified under "Posidonia bronni". Found usually associated with Driftwood.

Eolamprotula[3][4]

  • E. cremeri
  • E. sp.
  • Żarnów
  • Wąsosz
  • Kuraszków
  • Żarki 88-Z borehole
  • Choroń 31-BN borehole

Shells

A freshwater mussel, member of the family Unionidae inside Unionida. Associated with plant detritus, likely washed from nearby mainland

Lamprotula

Goniomya[18][20]

  • G. rhombifera
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

Shells

A Saltwater Clam, member of Pholadomyidae inside Pholadomyida. Rather common, but less abundant than other local genera

Gresslya[19]

  • G. sp.
  • Ahrensburg

Shells

A Saltwater Clam, member of Ceratomyidae inside Pleuromyoidea.

Lucina[18][20]

  • L. plana
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

Shells

A Saltwater Clam, member of Lucinidae inside Lucinida. Very abundant on the layers

Lucina

Meleagrinella[18][20][21]

  • M. substriata
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin clay pit
  • Gorzow Wielkopolski borehole
  • Kozłowice Clay Pit
  • "Boroszów" Brickyard
  • Żarki 88-Z borehole
  • Gorzków 43-BN borehole
  • Choroń 31-BN borehole
  • Bobolice 68-km borehole

Shells

A Saltwater Scallop, member of Oxytomidae inside Pectinida. Depositional environments in the Polish Basin at this time did not host organisms normally associated with well-oxygenated marine conditions. Meleagrinella substriata is the most common bivalve found on the Toarcian Polish Basin, linked with all the brackish Deposits.[21]

Meleagrinella

Nucula[9][22]

  • N. caecilia
  • Dobbertin

Shells

A Saltwater Clam, member of Nuculidae inside Nuculida.

Nucula

Parainoceramya[9][22][20][12][18]

  • P. dubia
  • P. amygdaloides
  • P. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade

Shells

A Saltwater/Brackish Clam, member of Inoceramidae inside Myalinida. Pseudomytiloides dubius is a possible junior synonym. This genus resembles the Modern Pearl Oysters, although it was more likely a clam. The specimens are rather complete. It is the most common bivalve found locally. Is also very numerous in young specimens

Parainoceramya

Parallelodon[19]

  • P. cypriniformis
  • Ahrensburg

Shells

A Saltwater Ark Clam, member of Parallelodontidae inside Arcida.

Stalagmina[20][18]

  • S. koeneni
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

Shells

A Saltwater Clam, member of Inoceramidae inside Myalinida. This genus resemble the Modern Pearl Oysters, although it was more likely a clam. The specimens are rather complete.

Steinmannia[17][18]

  • S. bronni
  • S. radiata
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin

Shells

A Saltwater Oyster, member of "Posidoniidae" inside Ostreoida. A genus classified under "Posidonia bronni".

Tancredia[19]

  • T. elegans
  • Ahrensburg

Shells

A Saltwater Clam, member of Tancrediidae inside Cardiida.

Gastropoda

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Genus Species Stratigraphic position Material Notes Images

Cerithium[18]

  • C. armatum
  • C. roeceri
  • Klein Lehmhagen pit, Grimmen

Shells

A Sea Snail, member of Cerithiinae inside Caenogastropoda.

Cerithium

Coelodiscus[9][22][20][19]

  • C. minutus
  • Clay pit of Dobbertin
  • Klein Lehmhagen pit, Grimmen
  • Müssentin gravel pit
  • Ahrensburg

Shells

A holoplanktonic Snail, type member of the family Coelodiscidae inside Prosobranchia. This is the oldest known holoplanktonic gastropod, thanks to bilateral symmetrical shells as an adaption to active swimming. Also the most common of the sea snails on the Clay Pit of Dobbertin, where is one of the most varied in size terms, with some of the biggest specimens of this snail from the Lower Toarcian know. It has been related to large floating driftwood as one of the primary settlers.

Turritella[22][20]

  • T. sp.
  • Dobbertin

Shells

A Sea Snail, member of Turritellidae inside Cerithioidea.

Cephalopoda

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Genus Species Stratigraphic position Material Notes Images

Acrocoelites[18]

  • A. pyramidalis
  • Klein Lehmhagen pit, Grimmen
  • Müssentin gravel pit

Multiple Specimens.

A Belemnoidean. member of the family Megateuthididae inside Belemnitida.

Acrocoelites

Beloteuthis[18]

  • B. schubleri
  • Klein Lehmhagen pit, Grimmen

Multiple Specimens

A Mesoteuthoidean, member of the family Beloteuthidae. Is a relatively small genus.

Beloteuthis

Dactylioceras[23][24][19]

  • D. athleticum
  • D. temperatum
  • D. attenuatum
  • D. tenuicostatum
  • D. semicelatum
  • D. cf. aequistriatum
  • D. eickenbergi
  • D. ernsti
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

Shells

An Ammonite, type member of the family Dactylioceratinae inside Ammonoidea. Most common ammonite found on the Green Series and the different erratic boulders, as is the most common on the German realm, on the north and the south, with several specimens of different sizes.

Dactylioceras

Eleganticeras[12][17][25][23][19]

  • E. elegantulum
  • E. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

Shells

An Ammonite, member of the family Hildoceratidae inside Ammonoidea.

Eleganticeras

Hildaites[19]

  • H. subserpentinus
  • Ahrensburg

Shells

An Ammonite, member of the family Hildoceratidae inside Ammonoidea.

Jeletzkyteuthis[18]

  • J. coriaceus
  • Klein Lehmhagen pit, Grimmen

Two Specimens

A Loligosepiidae Loligosepiidan (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. Gladii of Loligosepia can be distinguished from Jeletzkyteuthis by the transition lateral field/hyperbolar zone. Described originally as Belopeltis bollensis.

Lobolytoceras[12][25][23]

  • L. siemensi
  • L. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Thüringen
  • Brandenburg
  • Braunschweig
  • Boulder from Ahrensburg

Shells

An Ammonite, member of the family Lytoceratidae inside Ammonoidea.

Lobolytoceras

Nodicoeloceras[19]

  • N. crassoides
  • Ahrensburg

Shells

An Ammonite, type member of the family Dactylioceratinae inside Ammonoidea.

Peronoceras[12][25][19]

  • P. fibulatum
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

Shells

An Ammonite, member of the family Dactylioceratinae inside Ammonoidea.

Pseudolioceras[12][25][19]

  • P. lythense
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

Shells

An Ammonite, member of the family Harpoceratinae inside Ammonoidea.

Pseudolioceras

Phylloceras[19]

  • P. heterophyllum
  • Ahrensburg

Shells

An Ammonite, type member of the family Phylloceratinae inside Ammonoidea.

Dactylioceras

Teudopsis[18][26]

  • T. bollensis
  • Klein Lehmhagen pit, Grimmen

Few specimens

A Vampire Squid, member of Teudopsidae inside Vampyropodan. Related to the modern Vampyroteuthis infernalis.

Teudopsis

Tiltoniceras[12][17][25][23][19]

  • T. antiquum
  • T. costatum
  • T. acutum
  • T. schroederi
  • T. costatum
  • T. capillatum
  • T. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg

Shells

An Ammonite, member of the family Hildoceratidae inside Ammonoidea.

Tiltoniceras

Whitbyiceras[19]

  • W. pingue
  • Ahrensburg

Shells

An Ammonite, member of the family Phymatoceratidae inside Ammonoidea.

Crustacea

[edit]

Small indeterminate shrimps, sometimes found associated in great numbers, are recovered on several layers at Grimmen.[27]

Genus Species Stratigraphic position Material Notes Images

Bairdia[6]

  • B. hilda
  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Bairdiidae inside Bairdioidea.

Bythocypris[6]

  • B. elongata
  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Bythocyprididae inside Bairdioidea.

Cytherelloidea[6]

Indeterminate

  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Cytherelloidea inside Platycopina.

Euestheria[4][9][22][20][28][29][30][31]

  • E. opalina
  • E. loczyi
  • E. brodieana
  • E. minuta
  • Euestheria sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Lubienia
  • Kozlowice
  • Boroszów
  • Żuki
  • Gorzków
  • Choroń
  • Bobolice
  • Żarnów
  • Warsaw
  • Płońsk
  • Boża Wola
  • Lidzbark Warmiński
  • Polessk
  • Zakościele
  • Dąbrówka
  • Zakościtel
  • Sielec
  • Studzianna
  • Przysucha
  • Wąsosz
  • Kuraszków

Valves

A Freshwater Clam shrimp (Phyllopodan) of the family Lioestheriidae. Originally identified as a bivalve of the genus "Posidonia" and latter a member of the genus Lioestheria. They fed on detritus, being very small slow moving, nektonic organisms that filter fed as they floated. Euestheria opalina dominates the invertebrate assamblages. The specimens are numerous here and form thin layers, being the most abundant invertebrate recovered on all the formation. The presence of Euestheria marks the appearance of less saline conditions, as this is a mostly freshwater genus. Is linked with beds where abundant freshwater algae covers the layers, being both probably washed from the nearby mainland trought streams. The local Phyllopods are related with a great amount of freshwater debris (especially plants), and suggest seasonal changes on the rivers on the Toarcian Polish Basin. On Kozłowice, there is an association of the ichnogenus Planolites and phyllopods of this genus, which reflects a gradual decline in water salinity during a sea level high standard regression.

Glyphea[9][22][20]

  • G. liasina
  • Dobbertin
  • Isolated Molts
  • Isolated Chelae

A Marine Decapodan, type member of the family Glypheidae inside Decapoda.

Glyphea

Hungarella[6]

  • H. adenticulata
  • H. sp.
  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Healdiidae inside Metacopa. Hungarella adenticulata was first identified in Dobbertin

Liasina[3][13]

  • L. lanceolata
  • L. vestibulifera
  • Żuki
  • Gorzków
  • Choroń
  • Bobolice

Valves

A Marine Ostracodan of the family Pontocyprididae. Small marine ostracods related with abundant Green Algae environments

Paracypris[6]

  • P. semidisca
  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Paracypridinae inside Candonidae.

Polycope[6]

  • P. cincinnata
  • P. sp.
  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Polycopidae inside Cladocopina.

Procytheridea[6]

  • P. sp.
  • Dobbertin

Valves

A marine/brackish Ostracodan of the family Protocytheridae inside Cytheroidea.

Proeryon[22][27]

  • P. hartmanni
  • cf.P. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Isolated molts
  • Single specimen with preserved upper thorax and chelae

A Marine Decapodan, member of the family Proeryoninae inside Polychelida. Resemble Proeryon hartmani show less adaptations to hunt for small nectobenthic preys than other relatives, being abundant on Oyster-filled waters. There is a relative abundance of the genus in deep-water settings from the Toarcian onward.

Proeryon

Trachycythere[6]

  • T. tubulosa
  • Dobbertin

Valves

A marine/brackish Ostracodan of incertae sedis affinity

Arachnida

[edit]
Genus Species Stratigraphic position Material Notes Images

Seppo[32]

  • S. koponeni
  • Klein Lehmhagen pit, Grimmen

Single Incomplete Specimen

A spider, possible member of the superfamily Palpimanoidea.[32] It is the first confirmed spider from the lower Jurassic, and a rare find, probably washed to the sea due to a hard wind related to hurricane action. Probably a ground-dwelling predator that hunted the abundant insect fauna present on the layers.[32] With a robust and well-armed legs I, directed forwards give the suggestion that they were preycapture appendages, a morphology typical of a sit-and-wait predator, while the short legs III are more typical on web spiders, especially Orbweavers, but also found on Palpimanoids, but not on that that are common substrate dwellers, that had legs more equal.[32] Seppo was probably not a habitual ground dweller, with armoured front legs related to capturing dangerous prey, such as many palpimanoids today are Araneophagous, for example.[32]

Insecta

[edit]

Insects are common terrestrial animals that were probably drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation.[33] In Klein Lehmhagen insects are found as part of calcareous nodules in the exaratum-elegantulum subzones, with specimens also found in living chambers of Eleganticeras elegantulum macrochonchs and in fish coprolites which are the most frequent fossils at all.[33] In the elegantulum the insect fauna is dominated by beetle elytra, indicating strong fluvial input and a nearshore deltaic complex.[33] On Dobbertin, insects are present in the exaratum nodules, where fluvial input is seen thanks to the phyllopod abundance and whole bedding planes covered by algae substituted by Ca-phosphat, being the layers where insects are most abundant.[33]

Paraneoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Archipsylla[34][35]

  • A. primitiva
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Paraneopteran, member of the family Archipsyllidae inside Permopsocida.

Liassopsocus[35]

  • L. lanceolatus
  • Klein Lehmhagen pit, Grimmen

Specimens

A Paraneopteran, member of the family Psocidiidae inside Permopsocida.

Obotritagrion[35][36]

  • O. tenuiformum
  • O. petersi
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Complete Specimens
  • Isolated Wings

Primitive Pterygotans, member of the family Protomyrmeleontidae inside Dicondylia.

Protomyrmeleon[35][36]

  • P. brunonis
  • P. quadriordinum
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Prosepididontus[37][38]

  • P. calopteryx
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Reculidans, member of Gryllones (Extinct clade of Basal Insects) of the family Geinitziidae.

Grylloblattodea

[edit]
Genus Species Stratigraphic position Material Notes Images

Geinitzia[34][37]

  • G. schlieffeni
  • G. debilis
  • G. minor
  • Dobbertin, Mecklenburg (Greifswald collection)

Partial Specimens

An Ice crawler, member of the family Geinitziidae inside Grylloblattodea or Reculida.

Nele[35]

  • N. jurassica
  • Klein Lehmhagen pit, Grimmen

Partial Specimens

An Ice crawler, member of the family Bajanzhargalanidae inside Grylloblattodea. One of the main Floor insects found.

Grylloblatta, extant ice crawler, Nele was probably similar

Eoblattida

[edit]
Genus Species Stratigraphic position Material Notes Images

Griphopteron[39][35]

  • G. molle
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

An Eoblattidan, member of the family Blattogryllidae inside Eoblattida.

Thysanoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Liadoptilia[39]

  • L. misera
  • Klein Lehmhagen pit, Grimmen

Specimens

Thysanopterans, members of the family Lophioneurida inside Thripida. Aeroplankton is extraordinarily well preserved in Grimmen, with the most abundant representatives of the aeroplankton (around 3 mm) being Lophioneurids, specially Undacypha europaea.

Extant Thysanopteran, extinct genera were probably similar

Trichorthophlebia[39]

  • T. pilifera
  • Dobbertin, Mecklenburg (Greifswald collection)

Undacypha[35]

  • U. europaea
  • Dobbertin, Mecklenburg (Greifswald collection)

Odonata

[edit]
Genus Species Stratigraphic position Material Notes Images

Anisozygopteron[34]

  • A. geinitzianum
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Dragonfly, member of the family Myopophlebiidae inside Odonata.

Anomothemis[34]

  • A. brevistigma
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A dragonfly, incertae sedis inside Odonata.

Archithemis[22][34]

  • A. brodiei
  • Dobbertin, Mecklenburg (Greifswald collection)

Dialothemis[39]

  • D. dubia
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A Dragonfly, member of the family Selenothemistidae inside Odonata.

Eosagrion[37]

  • E. risi
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Dragonfly, member of the family Eosagrionidae inside Odonata.

Grimmenopteron[35]

  • G. elegantulum
  • Klein Lehmhagen pit, Grimmen

Specimens

A Dragonfly, member of the family Liassophlebiidae inside Odonata.

Heterophlebia[35][40]

  • H. buckmani
  • H. megapolitana
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)

Specimens

A Dragonfly, member of the family Heterophlebiidae inside Odonata.

Heterothemis[41][42]

  • H. brodiei
  • H. germanica
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Ahrensburg erratics
  • Isolated Wings

A Dragonfly, member of the family Liassogomphidae inside Odonata.

Liadothemis[34][39][40]

  • L. hydrodictyon
  • L. major
  • L. geinitzi
  • L. insignis
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A dragonfly, incertae sedis inside Odonata. Magnasupplephlebia represents a very large dragonfly, with a wingspan of 13 cm. Other odonatan remains are unable to being referred to a concrete group due to their incomplete status.

Magnasupplephlebia[40][43]

  • M. kallweita
  • M. intercalaria
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

Parelthothemis[34]

  • P. dobbertinensis
  • Dobbertin, Mecklenburg (Greifswald collection)

Petrothemis[34]

  • P. singularis
  • Dobbertin, Mecklenburg (Greifswald collection)

Pycnothemis[39]

  • P. densa
  • Dobbertin, Mecklenburg (Greifswald collection)

Rhabdothemis[39]

  • R. strigivena
  • Dobbertin, Mecklenburg (Greifswald collection)

Selenothemis[37]

  • S. liadis
  • Dobbertin, Mecklenburg (NHMW collection)
  • Isolated Wings

A Dragonfly, type member of the family Selenothemistidae inside Odonata.

Temnostigma[39]

  • T. singulare
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A dragonfly, incertae sedis inside Odonata. Magnasupplephlebia represents a very large dragonfly, with a wingspan of 13 cm. Other odonatan remains are unable to being referred to a concrete group due to their incomplete status.

Trigonophlebia[35]

  • T. zessini
  • Klein Lehmhagen pit, Grimmen

Specimens

A Dragonfly, member of the family Heterophlebiidae inside Odonata.

Turanopteron[35]

  • T. pommerana
  • Klein Lehmhagen pit, Grimmen

Specimens

A Dragonfly, member of the family Asiopteridae inside Odonata. "Sphenophlebia pommerana" is a junior synonym.

Plecoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Dicronemoura[35]

  • D. furcata
  • Klein Lehmhagen pit, Grimmen

Specimens

A stonefly, member of the family Perlariopseidae inside Plecoptera.

Dobbertiniopteryx[44]

  • D. capniomimus
  • Dobbertin, Mecklenburg (Ansorge coll)

Specimens

A small winter Stonefly, member of the family Capniidae inside Plecoptera.

extant Capniidae, Dobbertiniopteryx was probably similar

Orthoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Locustopsis[9][35][34][39][45][46]

  • L. gyra
  • L. bernstorffi
  • L. sippeli
  • L. dubia
  • L. bernstorffi
  • L. nana
  • L. reducta
  • L. elongata
  • L. elegans
  • L. pulchella
  • L. procera
  • L. mecklenburgica
  • L. maculosa
  • L. sp 1
  • L. sp 2
  • L. sp 3
  • L. sp 4
  • Grimmen (Krempien collection)
  • Lehmhagen, near Grimmen (Zessin collection)
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Complete Specimens
  • Isolated Wings

Grasshoppers of the family Locustopsidae inside Orthoptera.

Locustophanes[39]

  • L. rhipidophorus
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Locust, type member of the family Locustidae inside Orthoptera.

Acanthacris, extant Locust, Locustophanes was probably similar

Panorpidium[9][35][34][39][37][47]

  • P. geinitzi
  • P. magna
  • P. minima
  • P. media
  • P. reticulata
  • P. mesostena
  • Grimmen (Krempien collection)
  • Lehmhagen, near Grimmen (Zessin collection)
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Breslau Museum Coll.)
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Specimens
  • Isolated Wings

A Grasshopper, member of the family Elcanidae inside Orthoptera. The species Panorpidium magna is among the largest Orthopterans of the Jurassic, while Panorpidium minima is among the smallest.

Parelcana[35][39][47]

  • P. tenuis
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)

Plesioschwinzia[48]

  • P. thalassophila
  • P. lacera
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Grasshoppers of the family Locustopsidae inside Orthoptera.

Protochaeta[35][39]

  • P. masculina
  • P. lanceolata
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Grasshopper, member of the family Regiatidae inside Orthoptera.

Protogryllus[35][34][37][39][42][49]

  • P. dobbertinensis
  • P. germanicus
  • P. sp.
  • P. vicinus
  • P. liadis
  • P. major
  • P. stenobasis
  • P. femina
  • P. acutipennis
  • P. minor
  • P. irregularis
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (BMNH collection)
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Ahrensburg Erratics

Specimens

A Cricket-like Grasshopper, type member of the family Protogryllidae inside Orthoptera.

Schwinzia[45]

  • S. sola
  • Dobbertin, Mecklenburg (Zessin collection)
  • Isolated Wings

Grasshoppers of the family Locustopsidae inside Orthoptera.

Synelcana[20][48]

  • S. muelleri
  • Dobbertin, Mecklenburg (Zessin collection)
  • Isolated Wings

A Grasshopper, member of the family Elcanidae inside Orthoptera.

Zalmonites[34]

  • Z. geinitzi
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A grasshopper, incertae Sedis inside Orthoptera.

Phasmatodea

[edit]
Genus Species Stratigraphic position Material Notes Images

Durnovaria[35]

  • D. cf. parallela
  • Klein Lehmhagen pit, Grimmen

Specimens

A Stick Insect, member of the family Aerophasmidae inside Phasmatodea.

Example of extant Phasmatodean, Durnovaria was probably similar

Blattodea

[edit]
Genus Species Stratigraphic position Material Notes Images

Blattula[22][39][41][50][42]

  • B. langfeldti
  • B. dubia
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Ahrensburg Erratics
  • Isolated Wings

A Cockroach, member of the family Blattulidae inside Dicondylia. Member of an extinct cockroach-like family which was widely distributed around the world and lasted from Late Triassic to Cretaceous.

Caloblattina[34][50]

  • C. mathildae
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Cockroach, type member of the family Caloblattinidae inside Blattodea.

Dipluroblattina[22][39]

  • D. scudderi
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A Cockroach, member of the family Blattulidae inside Dicondylia. Member of an extinct cockroach-like family which was widely distributed around the world and lasted from Late Triassic to Cretaceous.

Eublattula[39]

  • E. crassivena
  • Dobbertin, Mecklenburg (Greifswald collection)

Liadoblattina[39][41][50]

  • L. blakei
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Carnivorous Cockroach, type member of the family Raphidiomimidae inside Blattodea.

Mesoblattina[9][50]

  • M. protypa
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Cockroach, type member of the family Mesoblattinidae inside Blattodea.

Hemiptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Acromocoris[51]

  • A. similis
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A water boatman, member of the family Corixidae inside Hemiptera.

Example of extant Corixidae, Acromocoris was probably similar

Anosmus[39]

  • A. spilopterus
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Shore bugs, member of the family Archegocimicidae inside Hemiptera.

Aphlebocoris[34][41]

  • A. punctata
  • A. nana
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A creeping water bug, member of the family Naucoridae inside Hemiptera.

Example of extant Naucoridae, extinct genera where probably similar

Apopnus[41]

  • A. magniclavus
  • Dobbertin, Mecklenburg (Greifswald collection)

Apsicoria[39]

  • Apsicoria semideleta
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

True Bugs, member of the family Pachymeridiidae inside Hemiptera.

Ardela[35]

  • A. grimmenensis
  • Klein Lehmhagen pit, Grimmen

Specimens

Leafhoppers of the family Archijassidae inside Hemiptera.

Archegocimex[37][52]

  • A. geinitzi
  • A. primitiva
  • A. liadis
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Shore bugs, member of the family Archegocimicidae inside Hemiptera.

Example of extant Pentacora, Archegocimicidae genera where probably similar

Archicercopis[39]

  • A. falcatus
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Moss bugs of the family Progonocimicidae inside Hemiptera.

Archijassus[34][35][39]

  • Archijassus heeri
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Leafhoppers of the family Archijassidae inside Hemiptera.

Example of extant Membracidae, Archijassidae genera where probably similar

Aphidulum[35]

  • A. ciliatum
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Sternorrhynchan, member of the family Protopsyllidiidae inside Hemiptera.

Archiconiopteryx[35][34]

  • A. liasina
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Stemorrhynchan, type member of the family Archiconiopterygidae inside Hemiptera.

Cathalus[39]

  • Cathalus alutaceus
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

True Bugs, member of the family Pachymeridiidae inside Hemiptera.

Cixiites[34]

  • C. liasinus
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Planthoppers of the family Fulgoridiidae inside Hemiptera.

Cuneocoris[37]

  • C. geinitzi
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A true Bug, member of the family Cuneocoridae inside Hemiptera.

Diatillus[41]

  • Diatillus debilis
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Shore bugs, member of the family Archegocimicidae inside Hemiptera.

Eocercopis[39]

  • E. ancyloptera
  • E. similis
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Moss bugs of the family Progonocimicidae inside Hemiptera.

Fulgoridium[35][34][39]

  • F. egens
  • F. lapideum
  • F. dubium
  • F. latum
  • F. venosum
  • F. pallidum
  • F. geinitzi
  • F. spilographum
  • F. oligoneurum
  • F. regulare
  • F. graphipterum
  • F. inconspicuum
  • F. quadrisignatum
  • F. breviradiatum
  • F. modestum
  • F. anale
  • F. marginepunctatum
  • F. picturatum
  • F. anomalum
  • F. stigmaticum
  • F. punctatum
  • F. mortuum
  • F. alatum
  • F. nodosum
  • F. megapolitanum
  • F. pulchrum
  • F. remotum
  • F. multivenosum
  • F. debile
  • F. trifurcatum
  • F. litorale
  • F. parvispilum
  • F. retractum
  • F. polyneurum
  • F. oligospilum
  • F. nubeculum
  • F. obtusum
  • F. grave
  • F. fenestratum
  • F. breve
  • F. dilutum
  • F. multipunctatum
  • F. ampliatum
  • F. clavatum
  • F. nebulosum
  • F. vicinum
  • F. curvipenne
  • F. elegantulum
  • F. defunctum
  • F. plicatum
  • F. exhumatum
  • F. angulosum
  • F. acutum
  • F. rotundatum
  • F. ancylla
  • F. bifurcatum
  • F. inaequale
  • F. brachyptilum
  • F. intercalatum
  • F. balticum
  • Grimmen (EMAU collection)
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Breslau Museum Coll.)
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Complete Specimens
  • Partial Specimens
  • Isolated Wings

Planthoppers of the family Fulgoridiidae inside Hemiptera. The colossal abundance of the genus Fulgoridium maybe it is related to a preference for seashore habitats. Some specimens are indistinguishable, making possible some species synonymous.

Example of extant Fulgoridae, Fulgoridiidae genera where probably similar

Grimmenaphis[35]

  • G. magnifica
  • Klein Lehmhagen pit, Grimmen

Specimens

An aphid, member of the family Oviparosiphidae inside Hemiptera. Among the oldest Aphids

Example of extant Aphidoidea, Oviparosiphidae genera where probably similar

Hadrocoris[39]

  • H. scutellaris
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A true Bug, member of the family Hadrocoridae inside Hemiptera.

Hypocimex[39]

  • Hypocimex membranaceus
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

True Bugs, member of the family Pachymeridiidae inside Hemiptera.

Liadopsylla[35][41]

  • L. geinitzi
  • L. obtusa
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A jumping plant louse, member of the family Liadopsyllidae inside Hemiptera.

Liassocercopis[35]

  • L. schnicki
  • Klein Lehmhagen pit, Grimmen

Specimens

A cicadomorphan, member of the family Hylicellidae inside Hemiptera.

Liasocoris[53]

  • L. hainmuelleri
  • Schwinzer brickyard, near Goldberg

Specimens

A true bug, incertae sedis inside Hemiptera.

Margaroptilon[39]

  • M. germanicum
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Planthoppers of the family Fulgoridiidae inside Hemiptera.

Mesoledra[35][39]

  • M. pachyneura
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Naturkundemuseum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Leafhoppers of the family Archijassidae inside Hemiptera.

Metafulgoridium[39]

  • M. spilotum
  • M. ampliatum
  • M. graptum
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Planthoppers of the family Fulgoridiidae inside Hemiptera.

Pachymeridium[9]

  • P. dubium
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

True Bugs, member of the family Pachymeridiidae inside Hemiptera.

Probascanion[39][51]

  • P. megacephalum
  • Dobbertin, Mecklenburg (NHMW collection)

Specimens

A Coleorrhynchan, member of the family Probascanionidae inside Hemiptera.

Procercopis[22][34][39]

  • P. alutacea
  • P. coriacea
  • P. liasina
  • P. similis
  • P. jurassica
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Froghoppers of the family Procercopidae inside Hemiptera.

Procercopina[35]

  • P. frenzeli
  • Klein Lehmhagen pit, Grimmen

Progonocimex[37][52]

  • P. jurassicus
  • P. liasinus
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Moss bugs of the family Progonocimicidae inside Hemiptera.

Example of extant Peloridiidae, Progonocimicidae genera where probably similar

Progonocoris[37]

  • P. pictus
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

Shore bugs, member of the family Archegocimicidae inside Hemiptera.

Sisyrocoris[41]

  • S. rudis
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

True Bugs, member of the family Pachymeridiidae inside Hemiptera.

Toarcopsyllidium[35]

  • T. ernsti
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Sternorrhynchan, member of the family Protopsyllidiidae inside Hemiptera.

Hymenoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Brigittepterus[54]

  • B. brauckmanni
  • Dobbertin, Mecklenburg (Zessin collection)

Isolated Wings

Wasps, member of the family Ephialtitidae inside Hymenoptera

Grimmaratavites[55]

  • G. mirabilis
  • Klein Lehmhagen pit, Grimmen

Specimens

A Wood Wasp, member of the family Karatavitidae inside Hymenoptera.

Liadobracona[54]

  • L. raduhna
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

Isolated Wings

Wasps, member of the family Ephialtitidae inside Hymenoptera

Xyelula[54]

  • X. benderi
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

Specimens

A pseudo-Wasp, member of the family Sepulcidae inside Hymenoptera.

Megaloptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Dobbertinia[37]

  • D. reticulata
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

An alderfly, member of the family Sialidae inside Megaloptera.

Example of extant Sialidae, Dobbertinia was probably similar

Neuroptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Actinophlebia[34][35][39]

  • A. megapolitana
  • A. aenea
  • A. parallela
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Lacewings of the family Prohemerobiidae inside Neuroptera.

Apeirophlebia[34]

  • A. grandis
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A silky lacewing, member of the Family Psychopsidae inside Neuroptera. An unexpected giant Silky lacewing, with a size up to 7 cm.

Example of extant Psychopsidae, Apeirophlebia was probably similar

Epigambria[39][56]

  • E. longipennis
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A lacewing, member of the Family Epigambriidae inside Neuroptera. Members of the macrofamily Ithonidae

Example of extant Ithonidae, Epigambriidae members where probably similar

Liassochrysa[57][58]

  • L. stigmatica
  • Dobbertin, Mecklenburg (Ansorge coll)

Specimens

A Neuropteran of the family Mantispidae. It is the earliest know Chrysopid

Example of extant Mantispidae, Liassochrysa was probably similar

Loxophleps[39]

  • L. costalis
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

A lacewing, incertae sedis inside Neuroptera. The wings if this neauropterans are rather fagmentary and its inclusion on any extant or extinct family is complex

Melaneimon[39]

  • M. dubium
  • Dobbertin, Mecklenburg (Greifswald collection)

Melamnous[39]

  • M. indistinctus
  • Dobbertin, Mecklenburg (Greifswald collection)

Mesoleon[34]

  • M. dobbertinianus
  • Dobbertin, Mecklenburg (Greifswald collection)

Mesosmylina[35]

  • M. falcifera
  • Klein Lehmhagen pit, Grimmen

Specimens

A lance lacewing, member of the family Osmylidae inside Neuroptera.

Polyosmylus[35][56]

  • P. excelsus
  • Klein Lehmhagen pit, Grimmen

Specimens

A lacewing, member of the Family Epigambriidae inside Neuroptera. Members of the macrofamily Ithonidae

Prohemerobius[34][35][39]

  • P. prodromus
  • P. ovatus
  • P. anomalus
  • P. costalis
  • P. persimilis
  • P. geinitzi
  • P. liasinus
  • P. major
  • P. parvulus
  • P. geinitzianus
  • P. dilaroides
  • P. chryseus
  • P. latus
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Lacewings of the family Prohemerobiidae inside Neuroptera.

Solenoptilon[59]

  • S. kochi
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A lacewing, type member of the Family Solenoptilidae inside Neuroptera.

Toarciconiopteryx[60]

  • T. dipterosimilis
  • Klein Lehmhagen pit, Grimmen

Isolated hind wing

A dustywing, member of the Family Toarciconiopteryginae inside Neuroptera. The oldest representative of the family

Example of extant Coniopterygidae, Toarciconiopteryx was probably similar

Coleoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Allognosis[61]

  • A. nitens
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera. Great amount of Beetle elytrons on both Dobbertin and Grimmen corroborates the presence of suitable environments nearby.

Anancaeon[39]

  • A. microcephalum
  • Dobbertin, Mecklenburg (NHMW collection)

Anomerus[39]

  • A. punctifer
  • Dobbertin, Mecklenburg (Greifswald collection)

Anypostatus[39]

  • A. taurus
  • Dobbertin, Mecklenburg (Greifswald collection)

Apioderes[39]

  • A. punctatus
  • Dobbertin, Mecklenburg (Greifswald collection)

Apsychus[39]

  • A. alutaceus
  • Dobbertin, Mecklenburg (Greifswald collection)

Bareus[39]

  • B. strigipennis
  • Dobbertin, Mecklenburg (Greifswald collection)

Bathygerus[61]

  • B. bellus
  • B. divergens
  • Dobbertin, Mecklenburg (Greifswald collection)

Carabites[34][61]

  • C. dubius
  • C. geinitzi
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A ground beetle, member of the family Carabidae inside Coleoptera.

Example of extant Carabidae, Carabites was probably similar

Clinomerus[39]

  • C. laticollis
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Coptogyrinus[34]

  • C. scutellatus
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A whirligig beetle, member of the family Gyrinidae inside Coleoptera.

Example of extant Gyrinidae, Coptogyrinus was probably similar

Ecthlimma[39]

  • E. forficuloides
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Enamma[34]

  • E. striatum
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Eurynucha[34]

  • E. pseudobuprestis
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

A jewel beetle, member of the family Buprestidae inside Coleoptera.

Example of extant Buprestidae, Eurynucha was probably similar

Gyrinulopsis[34]

  • G. nanu
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Hydrobiites[39]

  • H. dobbertinensis
  • H. punctatostriatus
  • Dobbertin, Mecklenburg (NHMW collection)

Elytrons

A beetle, member of the family Permosynidae inside Coleoptera.

Keleusticus[61]

  • K. zirkeli
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Liassodites[62]

  • L. obsti
  • Klein Lehmhagen pit, Grimmen
  • Single complete specimen

A False Ground Beetle of the family Trachypachidae.

Extant member of Trachypachidae, Liassodites was probably similar

Masselytron[39]

  • M. quinquestriatum
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Nannoodes[34]

  • N. pseudocistela
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Nebrioides[61]

  • N. dobbertinensis
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Onkedodimus[39]

  • O. discicollis
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Oxytoroptera[39]

  • O. mediocris
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Paracurculium[61]

  • P. punctatum
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Parnidium[34]

  • P. frechi
  • P. geinitzi
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Periboloptera[39]

  • P. rotunda
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Plastobuprestites[61]

  • P. elegans
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Plastonebria[61]

  • P. scudderi
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Polypamon[61]

  • P. byrrhoides
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Pseudocyphon[34]

  • P. geinitzi
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Pseudoprionites[61]

  • P. liasinus
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Elytrons

Beetles, incertae sedis inside Coleoptera.

Thoracotes[34]

  • T. dubius
  • Dobbertin, Mecklenburg (Greifswald collection)
  • 2 Complete Specimens
  • Isolated Elytrons

A bark gnawing beetle, member of the family Trogossitidae inside Coleoptera.

Example of extant Trogossitidae, Thoracotes was probably similar

Trichoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Archiptilia[39]

  • A. ovata
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Caddisflies, member of the family Necrotauliidae inside Trichoptera.

Epididontus[39]

  • E. geinitzianus
  • Dobbertin, Mecklenburg (Greifswald collection)

Liadotaulius[34][39][63]

  • L. maior
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A finger-net caddisfly, member of the family Philopotamidae inside Trichoptera.

Example of extant Philopotamidae, Liadotaulius was probably similar

Mesotrichopteridium[34][39][63]

  • M. intermedium
  • M. pusillum
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Caddisflies, member of the family Necrotauliidae inside Trichoptera.

Metarchitaulius[39]

  • M. longus
  • Dobbertin, Mecklenburg (Greifswald collection)

Nannotrichopteron[34]

  • N. gracile
  • Dobbertin, Mecklenburg (Greifswald collection)

Necrotaulius[39][63]

  • N. parvulus
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Palaeotaulius[39]

  • P. vicinus
  • Dobbertin, Mecklenburg (Greifswald collection)

Parataulius[39]

  • P. jurassicus
  • Dobbertin, Mecklenburg (Greifswald collection)

Paratrichopteridium[39]

  • P. efossum
  • P. costale
  • Dobbertin, Mecklenburg (Greifswald collection)

Pararchitaulius[39]

  • P. ovalis
  • Dobbertin, Mecklenburg (Greifswald collection)

Pseudorthophlebia[34]

  • P. platyptera
  • Dobbertin, Mecklenburg (Greifswald collection)

Lepidoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Eolepidopterigidae[64]

  • Eolepidopterigidae indet. A
  • Eolepidopterigidae indet. B
  • Eolepidopterigidae indet. C
  • Klein Lehmhagen pit, Grimmen
  • Isolated Wings
  • Isolated Wing Scales

A Moth, member of the family Eolepidopterigidae inside Lepidoptera. They are essential fossils for the Development of the color on Lepidopterans.

Mecoptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Mesobittacus[35]

  • M. minutus
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Hangingflies, member of the family Bittacidae inside Mecoptera.

Mesochorista[39]

  • M. sinuata
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Scorpionfly, member of the family Permochoristidae inside Mecoptera.

Mesopanorpa[39]

  • M. maculata
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Hangingflies, member of the family Orthophlebiidae inside Mecoptera.

Neorthophlebia[35][34][39]

  • N. maculipennis
  • N. megapolitana
  • N. pallida
  • N. debilis
  • N. simillima
  • N. medialis
  • N. stigmatica
  • N. acutipennis
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Hangingflies, member of the family Bittacidae inside Mecoptera.

Example of extant Bittacidae, extinct members where probably similar

Orthophlebia[22][34][39][35]

  • O. germanica
  • O. elongata
  • O. fuscipennis
  • O. limnophila
  • O. cf. limnophila
  • O. radialis
  • O. liadis
  • O. maculata
  • O. vicina
  • O. brunsvicensis
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin (Breslau Museum Coll.)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Hangingflies, member of the family Orthophlebiidae inside Mecoptera.

Parabittacus[65]

  • P. analis
  • P. lingula
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

Hangingflies, member of the family Bittacidae inside Mecoptera.

Pseudopolycentropus[22][35]

  • P. triangularis
  • P. perlaeformis
  • P. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A scorpionfly, type member of the family Pseudopolycentropodidae inside Mecoptera.

Diptera

[edit]
Genus Species Stratigraphic position Material Notes Images

Aenne[66]

  • A. liasina
  • Schönenwald railway cutting, Grimmen
  • Klein Lehmhagen pit, Grimmen
  • Isolated Wings

The oldest known non biting Midges, members of the Chironomidae inside Diptera

Example of extant Chironomidae, extinct members where probably similar

Antefungivora[35]

  • A. germanica
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

Flies, member of the family Antefungivoridae inside Diptera.

Archibio[39]

  • A. mycetophilinus
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Archipleciomima[35][67]

  • A. germanica
  • A. sp.
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A fly, incertae sedis inside Diptera

Archirhyphus[39]

  • A. geinitzi
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Flies, member of the family Protorhyphidae inside Diptera

Architipula[34][39][68]

  • A. stigmatica
  • A. latipennis
  • A. elegans
  • A. seebachi
  • A. nana
  • A. pusilla
  • A. pulla
  • A. debilis
  • A. simplex
  • A. intermedia
  • A. obliqua
  • A. areolata
  • A. maior
  • A. conspicua
  • A. clara
  • A. dubia
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A Crane Fly of the family Limoniidae inside Diptera.

Example of extant Limoniidae, extinct members where probably similar

Crenoptychoptera[69]

  • C. dobbertinensis
  • Dobbertin, Mecklenburg (Ansorge coll)

Isolated Wings

A Phantom crane fly, member of the family Ptychopteridae inside Diptera.

Example of extant Ptychopteridae, extinct members where probably similar

Eoditomyia[35]

  • E. primitiva
  • Klein Lehmhagen pit, Grimmen

Specimens

A Fly, type member of the family Eoditomyidae inside Diptera

Eolimnobia[34]

  • E. geinitzi
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

A Phantom crane fly, member of the family Ptychopteridae inside Diptera.

Example of extant Ptychopteridae, extinct members where probably similar

Eoptychoptera[35][34][39]

  • E. simplex
  • E. eximia
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Greifswald collection)

Eotipula[34]

  • E. defuncta
  • E. mortua
  • E. coarctata
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A Crane Fly of the family Limoniidae inside Diptera.

Grimmenia[68]

  • G. prima
  • G. secunda
  • Lehmhagen, near Grimmen (Zessin collection)
  • Isolated Wings

A Crane Fly of the family Limoniidae inside Diptera.

Grimmyia[35]

  • G. baltica
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Snipe Fly, member of the family Rhagionidae inside Diptera.

Example of extant Rhagionidae, extinct members where probably similar

Heterorhyphus[35][39]

  • H. triangularis
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Fly, type member of the family Heterorhyphidae inside Diptera

Liassobrachyceron[35][70]

  • L. kotejai
  • Dobbertin, Mecklenburg (Ansorge coll)

Isolated Wings

A Snipe Fly, member of the family Rhagionidae inside Diptera.

Liassopsychodina[71]

  • L. pommerana
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Moth fly, member of the family Psychodidae inside Diptera.

Example of extant Psychodidae, extinct members where probably similar

Mailotrichocera[72]

  • M. mikereichi
  • M. variabilis
  • M. zessini
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Ansorge coll)

Isolated wings

A Winter Crane Fly, member of the family Trichoceridae inside Diptera.

Example of extant Trichoceridae, extinct members where probably similar

Mesorhyphus[37][39]

  • M. nanus
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

A Wood Gnat, member of the family Anisopodidae inside Diptera.

Example of extant Anisopodidae, extinct members where probably similar

Mesotipula[37][35][34][39][68][73]

  • M. siggiae
  • M. vicina
  • M. cf. vicina
  • M. mirabilis
  • M. curvata
  • M. geinitzi
  • M. minuta
  • M. lapidaria
  • M. parva
  • Lehmhagen, near Grimmen (Zessin collection)
  • Dobbertin, Mecklenburg (NHMW collection)
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A Crane Fly of the family Limoniidae inside Diptera.

Metatrichopteridium[74]

  • M. confusum
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

A Fly, member of the family Hennigmatidae inside Diptera.

Nannotanyderus[35][71]

  • N. grimmenensis
  • N. krzeminskii
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

Isolated Wings

A primitive Crane Fly, member of the family Tanyderidae inside Diptera.

Oryctochlus[35][66]

  • O. toarciensis
  • Klein Lehmhagen pit, Grimmen
  • Isolated Wings

The oldest known non biting Midges, members of the Chironomidae inside Diptera

Rhaetomyia[35]

  • R. herrigi
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Phantom midge of the family Chaoboridae inside Diptera.

Example of extant Chaoboridae, extinct members where probably similar

Palaeobrachyceron[35]

  • P. willmanni
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Snipe Fly, member of the family Rhagionidae inside Diptera.

Phryganidium[22][39]

  • P. minimum
  • Dobbertin, Mecklenburg (Greifswald collection)
  • Isolated Wings

A Crane Fly of the family Limoniidae inside Diptera.

Pleciofungivora[35]

  • P. liassica
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Fly, member of the family Pleciofungivoridae inside Diptera.

Podonomius[35][66]

  • P. tumidus
  • Klein Lehmhagen pit, Grimmen
  • Isolated Wings

The oldest known non biting Midges, members of the Chironomidae inside Diptera

Praemacrochile[75]

  • P. decipiens
  • P. dobbertinensis
  • Dobbertin, Mecklenburg (Zessin collection)
  • Dobbertin, Mecklenburg (Ansorge coll)

Isolated Wings

A primitive Crane Fly, member of the family Tanyderidae inside Diptera.

Protobrachyceron[37]

  • P. liasinum
  • Dobbertin, Mecklenburg (Ansorge coll)
  • Dobbertin, Mecklenburg (Greifswald collection)

Specimens

A Fly, member of the family Protobrachyceridae inside Diptera

Protoplecia[22][35]

  • P. klafacki
  • P. liasina
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated wings

A Fly, type member of the family Protopleciidae inside Diptera.

Protorhyphus[35][39]

  • P. simplex
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, Mecklenburg (Greifswald collection)

Isolated Wings

Flies, member of the family Protorhyphidae inside Diptera

Tanypsycha[71]

  • T. connexa
  • Klein Lehmhagen pit, Grimmen

Isolated Wings

A Moth fly, member of the family Psychodidae inside Diptera.

Echinodermata

[edit]

In Dobbertin, the echinoderm remains are rare in contrast to foraminifera, phyllopods and ostracods, yet in some places they attain a percentage of the total fauna between 0.7 and 26.5%.[6] In the upper layers they're totally absent, as well on the erratics and in the whole Grimmen sequence.[76]

Genus Species Stratigraphic position Material Notes Images

Crinoidea[6]

Indeterminate

  • Dobbertin
  • Isolated Branches
  • Isolated Cirri
  • Isolated terminal claw of a pinnula

Indeterminate Crinoid remains. This group has very limited/scarce record on the facies

Holothuriida[6]

Indeterminate

  • Dobbertin
  • Isolated Hooks
  • Isolated Sclerites
  • Isolated Holothurian Wheels

A sea Cucumbers of uncertain placement. Remains of this group are very rare on the layers.

Sinosura[6]

  • S. brodiei
  • S. spp.
  • Dobbertin
  • Isolated Ambulacralia
  • Isolated Adambulacralia
  • Isolated Side shields
  • Isolated Ventral plates
  • Isolated Mouthpieces
  • Isolated Arm spines
  • Isolated Bursal braces

A Brittle Star, member of Ophioleucidae inside Ophioleucida. Identified as Ophiura longivertebralis. The majority of the Echinoderm material found belong to this genus and overall to Ophiuroids. This can be related to the fact some Ophiuroids are more tolerant with Brackish waters.

Vertebrates

[edit]

Fishes

[edit]

Actinopteri

[edit]
Genus Species Stratigraphic position Material Notes Images

Dapedium[22][76][77]

  • D. pholidotum
  • Dobbertin

Incomplete skull

A marine Osteichthyes, type member of the Dapediidae inside Neopterygii.

Dapedium

Grimmenichthys[78]

  • G. ansorgei
  • Cf. G. ansorgei
  • Klein Lehmhagen pit, Grimmen

Articulated, but incompletely preserved specimens

A marine Osteichthyes, member of the family Pholidophoriformes inside Teleostei. Found Exclusively on the Green Series, this genus was a small 'pholidophoriform' teleost. The total length remains unknown due to poor preservation, but probably does not exceed 100 mm.

Grimmenodon[76][79]

  • G. aureum
  • Klein Lehmhagen pit, Grimmen

Almost complete left prearticular with dentition

A marine Osteichthyes, Member of the order Pycnodontiformes inside Neopterygii. Is of comparatively small size, suggesting a hypothetical small standard length of the fish of about 7–10 cm at the time of death.[79] It is a taxon that shows clear adaptations to durophagy, inferring a predominant diet of especially hard food items.[79]

Gyrosteus[80]

  • G. mirabilis
  • Ahrensburg Erratic Assemblage

GPIH 4864, Hyomandibula

A marine Osteichthyes, member of the Chondrosteidae inside Acipenseriformes.

Lepidotes[80][76][81][82]

  • L. buelowianus
  • L. elvensis
  • L. spp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Ahrensburg Erratic Assemblage

Various specimens

A marine Osteichthyes, member of the family Lepidotidae inside Lepisosteiformes. Various species of the genus probably lived in the zone, yet the relationships of the fossils weren't studied.

Lepidotes

Lepisosteidae[78][83]

Indeterminate

  • Grimmen

GG 439/4-7 scales

The oldest confirmed true Gar, member of the Lepisosteidae inside Lepisosteiformes. The oldest reliable lepisosteiform remains.

Lepisosteus extant example of Gar

Leptolepidae[84]

Indeterminate

  • Former clay pit of Dobbertin
  • Isolated head with missing postcranial bones
  • Assigned Teeth

A marine Osteichthyes, member of the family Leptolepidae inside Teleostei.

Leptolepis[80][12][22][76][85][81][86][84]

  • L. bronni
  • L. coryphaenoides
  • L. normandica
  • L. jaegeri
  • L. spp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin clay pit
  • Karziger Ufer
  • Müssentin
  • Broock
  • Hohenbarnekow
  • Grenzin
  • Groß Roge
  • Elmenhorst
  • Klütz Höved
  • Pinnow
  • Consrade
  • Ahrensburg
  • Slightly disarticulated, incomplete head; Subadult, almost complete; fragmentary head; isolated head with pectoral girdle; incomplete head
  • Isolated remains in form of "Grätensandsteine"

A marine Osteichthyes, type member of the family Leptolepidae inside Teleostei. The most abundant vertebrate recovered on the formation, including 3D preserved specimens, as well, is the main component of the Fishbone sandstones from the upper layers.

Leptolepis

Lycopteroidarum[85]

  • L. sp. A
  • L. sp. B
  • Ahrensburg Erratic Assemblage
  • Otoliths

Brackish/Freshwater Osteichthyes otoliths of the family Lycopteroidea inside Lycopteriformes.

Mengius[87]
  • M. acutidens
  • Klein Lehmhagen pit, Grimmen
  • Schwinzer Ziegelei ca 3 km E of Dobbertin
  • Holotype: Articulated, incomplete fish
  • Referred articulated, semiarticulated and fragmentary specimens
A marine/Brackish Osteichthyes, member of the family Lepidotidae inside Lepisosteiformes. Stomach content is also preserved on a specimen from Dobbertin, and is composed by arthropod cuticles.[88]

Pachycormiformes[80][78]

Indeterminate

  • Klein Lehmhagen pit, Grimmen

Unknown, only cited.

A marine Osteichthyes, member of the family Pachycormiformes inside Teleostei. The first find of the group in the region.

Pholidophorus[80][85]

  • P. sp.
  • Ahrensburg Erratic Assemblage
  • Assigned Fragmentary remains

A marine Osteichthyes, member of the family Pholidophoriformes inside Teleostei.

Pholidophorus

Pholidophoriformes[78]

Indeterminate

  • Klein Lehmhagen pit, Grimmen
  • Isolated skull roof with articulated ethmoid region, parasphenoid, and partial braincase

A marine Osteichthyes, member of the family Pholidophoriformes inside Teleostei. Resmble Pholidophoroides crenulata and P. limbata.

Saurichthys?[14]

  • S. sp
  • Gorzów Wielkopolski IG 1 borehole
  • Teeth

A Freshwater/brackish/marine Osteichthyes, type member of the family Saurichthyidae inside Chondrostei. This genus is known mostly from the late Triassic deposits on Europe, making its appearance on Toarcian strata superficially dubious. It can be a member of the genus Saurorhynchus, although this last one has never been found on Brackish-Prodeltaic Strata.

Saurichthys

Saurorhynchus[76][89]

  • S. hauffi
  • S. sp.
  • Klein Lehmhagen pit, Grimmen
  • Dobbertin
  • Diganostic Incomplete skull
  • Incomplete skull

A marine/brackish Osteichthyes, the youngest representative of the family Saurichthyidae inside Chondrostei. Distinctive for its large jaws, similar to modern Belonidae, it is a genus known from several locations across Europe on the Lower-Middle Jurassic.

Saurorhynchus

Proleptolepis[76][84]

P. sp.

  • Klein Lehmhagen pit, Grimmen
  • MV 202612, fragmentary head .[84]
  • Assigned Teeth

A marine Osteichthyes, member of the family Leptolepidae inside Teleostei. Proleptolepis in the lower Toarcian of Grimmen suggests that this genus might have had a much wider palaeobiogeographical and temporal distribution, since specimens attributed to Proleptolepis have previously been reported only from the Sinemurian of western Europe.

Tetragonolepis[22][76]

  • T. semicincta
  • Klein Lehmhagen pit, Grimmen
  • Incomplete specimen
  • Isolated Jaw

A marine Osteichthyes, member of the family Semionotidae inside Semionotiformes. Dapedium and Tetragonolepis may have used alternative strategies in order to partition food resources efficiently and avoid direct competition.

Tetragonolepis

Chondrichthyes

[edit]
Genus Species Stratigraphic position Material Notes Images

Hybodus[76][85][80]

  • H. hauffianus
  • H. spp.
  • Klein Lehmhagen pit, Grimmen
  • Ahrensburg erratic assemblage
  • Gorzów Wielkopolski IG 1 borehole
  • Fragmentary Fin spine
  • Supposed coprolithes
  • Teeth

A marine Shark, member of the Hybodontiformes. Related to Hybodus hauffianus and other genera from the south of Germany. A Pliensbachian-aged deposit from Grimmen outcrop has revelated a pretty diverse Chondrichthyan fauna, including two species of the genus: Hybodus reticulatus & Hybodus hauffianus.[90]

Sarcopterygii

[edit]
Genus Species Location Material Notes Images

Ceratodus[14]

C. sp.

  • Gorzów Wielkopolski IG 1 borehole
  • Isolated tooth plates

A freshwater lungfish, type member of the Ceratodontidae inside Ceratodontiformes. Can be related to the Polish Triassic species Ceratodus silesiacus (Roemer, 1870), known from several isolated tooth plates found in the lacustrine Triassic horizon of Krasiejów. Several impressions of scales found in other boreholes can also belong to lungfish. The tooth plates were found at +819 m depth on Pomerania, on a layer with gray-green mudstone, lenticular lamination and siderite concretions. The abundance of megaspores indicate a deltaic or brackish environment.

Ceratodus reconstruction

Ichthyosaurs

[edit]
Genus Species Stratigraphic position Material Notes Images

Ichthyosauria[91][92][76][93]

Indeterminate

  • Klein Lehmhagen pit, Grimmen
  • Ahrensburg Erratic Assemblage
  • Presacral centrum
  • 2 ?sacral centrum
  • Isolated Vertebrae

An indeterminate ichthyosaur.

Neoichthyosauria[92][94]

Indeterminate

  • Clay Pit of Dobbertin

Four articulated tail vertebrae.

An indeterminate ichthyosaur, has been assigned to the species Stenopterygius longifrons.

Ophthalmosauridae[92][95][96][97]

Indeterminate

  • Klein Lehmhagen pit, Grimmen

Partial skull and associated postcranial elements preserved in a concretion

An indeterminate ichthyosaur. It has an expanded basipterygoid process on the basisphenoid, only currently known in members of the Ophthalmosauridae

Stenopterygius[92][76][98]

  • S. cf. quadriscissus
  • Clay pit of Dobbertin

Posterior left half of the cranium.

An Icthyosaur of the family Stenopterygiidae inside Thunnosauria. A common Toarcian Ichthyosaur, present on multiple layers. The rather exquisite level of preservation has led to know even the coloration.

Temnodontosaurus?[23][85]

  • T. cf. platyodon
  • Ahrensburg Erratic Assemblage
  • Skull
  • Coracoid
  • Associated rib fragment

An Icthyosaur, type member of Temnodontosauridae inside Neoichthyosauria. Assigned to Ichthyosaurus sp., but also suggested affinities to "Leptopterygius" (= Temnodontosaurus) platyodon.

Restoration

Sauropterygia

[edit]
Genus Species Stratigraphic position Material Notes Images

Meyerasaurus[91][23]

  • M. sp.
  • Ahrensburg Erratic Assemblage
  • Incomplete coracoid
  • Associated rib

A pliosauroid, member of the family Rhomaleosauridae inside Pliosauroidea.

Meyerasaurus

Plesiosauria[92][76]

Indeterminate

  • Klein Lehmhagen pit, Grimmen
  • Dobbertin, falciferum zone
  • Three caudal vertebrae and other postcranial elements
  • Gastralia
  • Phalanx
  • Cervical rib and other postcranial elements

A marine plesiosaur, member of the family Plesiosauria inside Sauropterygia. Non assigned to a concrete genus, probably are related to the Plesiosaurian genera from the Posidonia Shale

Plesiosauroidea[92][76]

Indeterminate

  • Klein Lehmhagen pit, Grimmen
  • Cervical centrum
  • Caudal centrum

A marine plesiosaur, member of the family Plesiosauria inside Sauropterygia. Non assigned to a concrete genus, probably are related to the Plesiosaurian genera from the Posidonia Shale

Seeleyosaurus[23][85]

  • S.? sp.
  • Ahrensburg Erratic Assemblage
  • Three articulated dorsal vertebrae

A plesiosaur, member of the family Plesiosauridae inside Plesiosauroidea. Related with the German Realm Fauna

Seeleysaurus

Crocodyliformes

[edit]
Genus Species Stratigraphic position Material Notes Images

Macrospondylus[92][76]

  • M. bollensis
  • Klein Lehmhagen pit, Grimmen

Incomplete skull and associated osteoderm

A marine Crocodrylomorph, member of the family Machimosauridae inside Teleosauroidea. A marine crocodylomorph with a diet probably based on fish. It was identified as Steneosaurus. Although highly incomplete, the preserved maximum antorbital width of about 50 mm demonstrates that GG 422/6 can be clearly referred to an immature individual.

Macrospondylus

Mesoeucrocodylia[92][76]

Indeterminate

  • Klein Lehmhagen pit, Grimmen

Cervical vertebra

A Crocodrilian, member of Mesoeucrocodylia. This cervical vertebra has resemblances with terrestrial suchians, such as Dianchungosaurus, but also cervicals of Sichuanosuchus. Is considered to come from an osteologically immature individual.

Sichuanosuchus

Mystriosaurus[92][76][93]

  • M. sp.
  • Klein Lehmhagen pit, Grimmen

Partial rostrum with teeth

A marine Crocodrylomorph, member of the family Teleosauridae inside Teleosauroidea. The tooth morphology and the internal rostral anatomy indicate that it can be positively referred to a longistrine thalattosuchian crocodyliform. The size of PLM uncatalogued, as compared to the corresponding portions of the rostra of other Toarcian thalattosuchians, is indicative of a juvenile individual, with expected around 200–250 mm long skull and the entire animal about 1.50 m.[93]

Teleosauridae[23][85][91]

Indeterminate

  • Ahrensburg Erratic Assemblage
  • Osteoderms
  • Isolated non described remains

A Crocodrylomorph, member of the family Thalattosuchia inside Neosuchia. Distinctive of the marine teleosaurid genus Macrospondylus.

Pterosauria

[edit]
Genus Species Stratigraphic position Material Notes Images

Pterosauria?[99]

Indeterminate
  • Klein Lehmhagen pit, Grimmen

GG 510, fragmentary tibiofibular(?)

A possible Pterosaur Bone, with features similar to the tibiofibular of coeval taxon Dorygnathus banthensis. Alternatively it can be another type of bone and belong to stem-turtles, plesiosaurs, dinosaurs or even anurans.[99]

Dorygnathus banthensis, know from inner localities, has shared features that may suggest GG 510 is from a Pterosaur

Dinosauria

[edit]
Genus Species Stratigraphic position Material Notes Images

Emausaurus[100][101]

  • E. ernsti
  • Klein Lehmhagen pit, Grimmen

Right side of the skull, the right lower jaw, caudal vertebrae, neural arches, a radius, a metatarsal, a claw, fragments of ribs, scutes and plates.[101]

An Ornithischian, basal member of Thyreophora inside Genasauria. Its juvenile status makes controversial its phylogeny, being considered a relative of the also Early Jurassic Genus Scelidosaurus.[101] The compacted pile of disarticulated cranial and postcranial elements of the basal thyreophoran Emausaurus has been suspected to be a Speiballen (i.e., a compacted mass of indigestible stomach contents) regurgitated by a large marine reptile.[101]

Emausaurus attacked by a theropod

Gravisauria[92][76][102][93][100]

Indeterminate

  • Klein Lehmhagen pit, Grimmen
  • Iliac preacetabular process
  • Distal right pubis
  • Proximal left ischium
  • Proximal right ischium
  • Single dorsal neural spine

A Saurischian, member of the clade Gravisauria inside Sauropoda. The specimen is believed to be a juvenile, based on the ossification and unfused spine. Has affinities with the genus Tazoudasaurus and it is clearly distinctive form the also Toarcian Ohmdenosaurus, who is thought to be more basal. The pelvic girdle elements can be clearly placed among the Sauropoda, on account of the presence of an elongated and strongly dorsally expanded iliac preacetabular process a possible relative. The ischia GG411/3-4 resemble those of Tazoudasaurus in exhibiting a subtriangular iliac peduncle which create a short anteriorly directed expansion to reach the medial acetabular rim, and are slightly less developed than the Genus Barapasaurus.

Barapasaurus can be a close relative of the Grimmen Sauropod

Megalosauridae[23][85][103]

Indeterminate

  • Forst Hagen gravel pit

Dorsal Vertebrae

A Saurischian, member of the family Megalosauridae inside Tetanurae. Was referred to Megalosaurus. The affinities of the Specimen aren't clear due to its fragmentary nature. Has been classified as Saurischia indeterminate, although shows clearly characters of the Orionides group (concave articular surfaces and a dished lateral pleurocoel, remnants of the neural arch and postzygapophyses).[103] The vertebrae centrum measures 80 mm, implying a medium-sized theropod (~5 m long). Can be related with Yunyangosaurus.

Marshosaurus, example of basal Orionides

Thyreophora[104]

Indeterminate

  • Klein Lehmhagen pit, Grimmen

GG 504, osteoderm

An Ornithischian, basal member of Thyreophora inside Genasauria. Interpreted as representing a lateral osteoderm of the neck or shoulder region of an early diverging thyreophoran

Plantae

[edit]

Coals

[edit]
A wildfire on a mountainside
Evidence of wildfires increases in the upper part of the formation.

The Łęka Coal Basin has been known since 1800 for its abundant deposits; younger material has been suggested as redeposited from the Paleozoic, but the basin primarily yields a series of Pliensbachian-Toarcian coals. In the Early Jurassic the Blanowice Formation was surrounded by land on the north, east and south, the source of sediment which was deposited as nearshore coal. This coal is abundant in the upper part of the formation, dominated by alluvial and lacustrine sand and coal-bearing sediments.[105] Organic matter associated with the coal includes the oldest known biomolecules (in the Mrzygłód clay-pit), composed of labdanoic acid, ferruginol, sugiol and 7-oxototarol.[105] The extracted samples were recovered at the Wysoka Lelowska 47Ż and Jaworznik 124Ż boreholes; five core samples were taken from the Żarki 90Ż core, the latter from an approximately 1.5-metre (4 ft 11 in) coal seam.[105] The random reflectance (%Rr) of the coals is 0.47–0.56, indicating sub-bituminous coal.[105] The geochemical data suggest low-maturity sediments,[105] and hopane isomers are relatively high in all samples.[105] This is confirmed by unstable biomolecules in the coal samples, including labdanoic acid, ferruginol, sugiol, oxototarol, beta-sitosterol and cholesterol.[105]

The coals are generally dominated by vitrinite macerals, except for a high percentage of inertinite. This indicates wildfires or peat fires, confirmed by charcoal fragments.[105] Sesquiterpenes and diterpenes, common in conifers, angiosperms and bryophytes, were also recovered from the coal.[106] Vitrinite has a reflectance value of 0.49-0.56 %Ro. The cupressaceae and podocarpaceae families are considered the main peat-forming plant species, due to the presence of phenolic abietanes and dehydroabietic acids.[105] Lignite indicates significant benzohopane derivatives in the surrounding sandstones, probable differences in biodegradation, and a typically low coalification range.[107] Later, larger studies note the influence of fires on the region.[108]

The Kaszewy coals, found in an approximately 150-metre (490 ft) section of terrestrial and marine siliciclastic sediments in the Kaszewy-1 and Niekłan PIG-1 boreholes, are the Ciechocinek Formation's major coals.[108] This section was in a nearshore-deltaic setting, with increased terrestrial and marine organic matter reflecting increased weathering and transport of terrestrial matter.[108] Abundant fossil charcoal and polycyclic aromatic hydrocarbons have been found. The number of coarse fossil charcoal particles (larger than 125 micrometers) in the Pliensbachian-Toarcian sections of the Kaszewy-1 core is very low (0-15 particles/10 g sediment), and fine charcoal particles (<125 μm) are more abundant (~12,000–256,000 particles/10 g sediment); there are also more non-charcoal particles.[108] There are more fine charcoal particles at the beginning of the Toarcian, reflecting environmental changes. In the polycyclic aromatic hydrocarbons, the pyrolytics (benz-anthracene, benzo(k)fluoranthene, fluoranthene, indeno[1,2,3-cd]pyrene, phenanthrene and pyrene) were detected in a wide variety of samples; phenanthrene is the most abundant component and coronene the least, suggesting the burning of organic matter.[108] Petrogenics are more abundant on the coal samples than pyrolytics, suggesting low wildfire activity.[108] Although the Kaszewy-1 borehole did not indicate increased wildfire activity, the fine fossil-charcoal abundance and pyrolytic concentration indicate regional wildfires.[108] Pyrolytics indicating the increased wildfire activity match the beginning of the Toarcian anoxic event, with intervals of fewer wildfires.[108] Wildfire changes match the Lower Toarcian negative carbon-isotopes emissions measured on the, which probably promoted a rise in atmospheric oxygen. Some questions remain; the climate was warmer and wetter (which can suppress wildfire activity), and wildfires persisted in the Kazewy-1 borehole wildfire activity was successfully sustained.[108] Wildfires may have subsided due to a lack of suitable fuel.[108]

Resins

[edit]
Type Location Material Notes Images

Fossil Resin[105]

  • Jaworznik 124Ż
  • Blanowice coals
  • Resinite
  • Resinous material

The local Resins containing sesqui-and diterpenoids that are produced by species of all contemporary conifer families, and in especially large amounts by Pinaceae, Cupressaceae, and Araucariaceae. The linked occurrence of inertinite, charcoal fragments, etc. show local Pliensbachian/Toarcian wildfires and/or peat fires.[105]

Extant Cupressaceae Resin, the Blanowice coals resins probably come from a similar Conifer

Floral Remains

[edit]

Polish Lower Toarcian palynology is assigned to the Paxillitriletes phyllicus (Ph) level (Isoetales), due to the abundance of this genus.[109] The lower part of the Toarcian level has numerous occurrences of this species, sometimes before the genera Erlansonisporites sparassis (Selaginella-like) and Minerisporites volucris (Isoetaceae) in the Gorzów Wlkp. IG 1 borehole.[109] The upper part contains less of the genus.[109] The most common species found on Poland in this era include Erlansonisporites sparassis, E. excavatus, Minerisporites volucris and Biharisporites scaber (Lycopodiopsida); Aneuletes potera and Trileites murrayi (both Selaginella) are found on the upper levels.[109] The Toarcian disturbance of the carbon cycle recorded on the Ciechocinek Formation, coincides roughly with the appearance of Paxillitriletes phyllicus. The type of dominant palynomorphs recovered changed from pollen grains during the Upper Pliensbachian to megaspores, indicating a climatic change from moderate and relatively dry to warm and humid in the early Toarcian.[109] This shift in local climate correlates with a global maritime transgression in which volcanism in the Karoo-Ferrar large igneous provinces raised the global temperature and disrupted the carbon cycle, creating a major greenhouse effect.[109] The prevalence of megaspore Paxillitriletes phyllicus correlate with warmth and humidity; the flora, dominated by the family Isoetaceae, requires standing water to reproduce.[109] The megaespore Paxillitriletes phyllicus then drops significantly, indicating a return to a more moderate climate during sedimentation of the younger Borucice Formation.[109]

The Blanowice Coals fossil wood from Zawiercie area were already described in 1917 as “Blanowicer Keuperholz”, on the basis of specimens from the “Elka”, “Kamilla” and “Zygmunt” coal pits, claimed to be xylologically similar, yet no taxon was named and the collection was not preserved.[110] Based on recent revisions of the local flora, likely belong to Agathoxylon.

The Lublin upland fluvial sandstones contain diverse fossil flora, associated genera and species with Lower Jurassic sediments. Carboniferous flora, similar to that of Jurassic formations, appeared in boreholes near the planned Bogdanka Coal Mine.[111] The age of the plant material was not determined until 2020, when it was identified as Lower Toarcian (with some Late Pliensbachian strata).[112] The Brody-Lubienia borehole is abundant in terrestrial palynomorphs and aquatic biomass.[112] The sediments from Brody-Lubienia have a more-terrestrial character, indicated by the frequent occurrence of plant roots and paleosol horizons;[112] moisture was probably fresh (not saline) water.[112] The environment was probably dry, developing flora near freshwater inflow from the east.[112]

The Lublin lias is dominated by cycads and Bennetites Ginkgoales. Ferns occur sporadically in the bottom of the Toarcian, where deposits are filled with coal, mudstone, sandstone and clay siderite (reworked from the Carboniferous) and pebbles from Devonian limestones.[113] Similar boreholes and nearby deposits indicate the end of a river which transported Devonian-Carboniferous deposits from the northeast to the aquatic inland environment.[113] Vegetation grew primarily outside the sedimentation area, on shores and in shallow water.[114]

At Ahrensburg there are plant remains in all horizons: wood, plant chaff and, in the Eleganticeras layers, nutty fruits, Araucariaceous cones, conifer branches, horsetail, etc.[19]

Bryophyta

[edit]
Genus Species Stratigraphic position Material Notes Images

Cingulatisporites[115]

  • C. floridus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Blanowice Coals
  • Spores

Affinities with Anthocerotaceae (specially Phaeoceros, Megaceros) inside Bryophyta. Moss spores from humid settings

Extant Phaeoceros, whose spores are similar to the extinct Cingulatisporites

Cirratriradites[115]

  • C. punctatus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Blanowice Coals
  • Spores
Affinities with Anthocerotaceae inside Bryophyta. Moss spores from humid settings
Extant Anthoceros, the typical example of the Anthocerotaceae. Cirratriradites probably come from a similar Plant

Foraminisporis[116]

  • F. dailyi
  • Brody-Lubienia borehole.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Spores

Rogalskaisporites[117][118]

  • R. cicatricosus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Blanowice Coals
  • Spores

Affinities with Sphagnopsida inside Sphagnales. Spores nearly identical to that one found associated with the modern moss genus Sphagnum. Moss related to high humid environments.

Extant Sphagnum, the typical example of the Sphagnales. Rogalskaiporites probably come from a similar Plant

Lycophyta

[edit]
Genus Species Stratigraphic position Material Notes Images

Acanthotriletes[119][116]

  • A. levidensis
  • A. minor
  • A. osmundiformis
  • A. rarispinosus
  • A. trigonis
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Spores

Affinities with the Selaginellaceae inside Lycopsida. Herbaceous Lycophyte flora, similar to Ferns, related with Humid Settings. This Family of Spores are also the most diverse on the Formation.

Aneuletes[120][121][116]

  • A. patera
  • Gorzów Wielkopolski IG 1
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Bacutriletes[120][122]

  • B. spicatus
  • B. clavatus
  • B.hamatus
  • B. tylotus
  • Helenow Beds
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Biharisporites[122]

  • B. scabrus
  • Mechowo borehole
  • Gorzow Wielkopolski borehole
  • Spores

Affinities with Lycopodiaceae inside Lycopodiopsida. Resemble spores of the modern genus Lycopodium. If it belongs to a similar genus, represent low herbaceous flora spores.

Carnisporites[120][121][116]

  • C. granulatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Echitriletes[123][121][122]

  • E. hispidus
  • E. sp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Eehitriletes[120][121]

  • E. hispidus
  • Brody-Lubienia borehole.
  • Gorzów Wielkopolski IG 1
  • Mechowo borehole
  • Parkoszowice borehole
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Erlansonisporites[124][125][122][116]

  • E. sparassis
  • E. excavatus
  • E. retieulatus
  • E. sparassis
  • E. reticulatus
  • E. tegimentus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds

spores

Affinities with the Selaginellaceae inside Lycopsida.

Foveosporites[125][119][116]

  • F. canalis
  • F. ovatus
  • F. cavernatus
  • F. polymicroforatus
  • F. foveoreticulatus
  • F. sp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with Lycopodiaceae inside Lycopodiopsida.

Horstisporites[125][116]

  • H.areolatus
  • H. planatus
  • H. puteus
  • H. foveatus
  • H. pustulatus
  • H. reticulatus
  • H. semireticulatus
  • H. sp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Lycopodiumsporites[119][125]

  • L. gristhorpensis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores

Affinities with Lycopodiaceae inside Lycopodiopsida.

Extant Lycopodium, typical example of Lycopodiaceae. Lycopodiumsporites probably come from a similar or a related Plant

Minerisporites[124][15][125]

  • M. institus
  • M. volucris
  • M. richardsoni
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Paxillitriletes[125][124]

  • P. phyllicus
  • Gorzów Wielkopolski IG 1 Borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Brody-Lubienia borehole
  • Spores

Affinities with the Isoetales inside Lycophyta. Spores related to modern Isoetes, representing Small plants related to water bodies. Paxillitriletes comprises the main Megaspore zonation of the Toarcian of Poland, being the most abundant spore found on the Ciechocinek Formation, and the genus that marks the start of the strata. The Abundance of this genus along with Minerisporites institus points to very humid conditions during almost the entire T-OAE, upwards from CIE step 2 of the carbon isotope curves.

Extant Isoetes, typical example of Isoetales. Paxillitriletes probably come from a similar or a related Plant

Striatriletes[121][125]

  • S. reticulatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Trileites[125][15][121]

  • T. turbanaeformis
  • T. ales
  • T. mesozoicus
  • T. murray
  • T. calvus
  • T. candoris
  • T. pyramidalis
  • T. solitus
  • T. usitatus
  • T. pinguis
  • T. utilis
  • T. aerolatus
  • T. phyllicus
  • T. reticulatus
  • T. usitatus
  • Trileites spp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Selaginellaceae inside Lycopsida.

Triletes[125][15][121]

  • T. reticulatus
  • T. datura
  • T. litchi
  • T. phyllicus
  • T. retiarius
  • T. richardsoni
  • T. sparassis
  • T. utilis
  • T. wicheri
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores

Equisetidae

[edit]
Genus Species Stratigraphic position Material Notes Images

Florinisporites[115]

  • F. ovatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores

Affinities with Equisetopsida inside Equisetopsida. Found endemically on the Polish basin and adjacent areas.

Extant Equisetum cone, the typical example of the Equisetopsida. Florinisporites spores are pretty similar to the extant ones

Irrimales[115]

  • I. laevis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores

Filicopsida

[edit]
Genus Species Stratigraphic position Material Notes Images

Baculatisporites[115]

  • B. fuscus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals

Spores

Affinities with the family Osmundaceae inside Polypodiopsida. Near Fluvial currents ferns, reted to the modern Osmunda Regalis.
Example of extant Osmunda specimens, Baculatisporites and Todisporites come probably from similar genera or maybe a species from the genus

Cardioangulina[116]

  • C. parva
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Spores
Affinities with the Matoniaceae inside Polypodiidae.

Corrugatisporites[120][121][116]

  • C. amplectiformis
  • C. scanicus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Schizaeaceae inside Schizaeales. Represents large spores from herbaceous ferns, that are more abundant on the south of the Toarcian polish basin than on the north.
Cyathidites[117][118][125][116]
  • C. minor
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals
Spores Affinities with Cyatheaceae inside Cyatheales. Cyathidites minor almost certainly belong to well known Mesozoic species Coniopteris hymenophylloides and to other fossil cyatheaceous or dicksoniaceous ferns such as Eboracia lobifolia and Dicksonia mariopteri.
Extant Cyathea, typical example of the Cyatheaceae. Cyathidites may have come from a similar genus
Crybelosporites[120][121][125][116]
  • C. stylosus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Marsileaceae inside Salviniales. Represents spore from fully aquatic ferns, found associated with fluvial or deltaic deposits, where probably formed large underwater colonies.
Extant Marsilea, typical example of the Marsileaceae. Crybelosporites may have come from a similar genus

Dennstaedtiites[115]

  • D. confragosus
  • D. concava
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals

Spores

Affinities with Dennstaedtiaceae inside Pteridophyta. Floor Herbaceous ferns, probably from covered forest areas.
Extant Pteridium aquilinum, typical example of the Dennstaedtiaceae. Dennstaedtiites spores are pretty similar to the extant ones of this genus
Gleicheniidites[120][121][125][116]
  • G. carinatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Gleicheniaceae inside Polypodiidae. Resemble the modern Gleichenia Spores, and probably represent a similar genus or a member of it. Fern related to large colonies, found mostly on humid environments.
Extant Unidentified Gleicheniaceae. Gleicheniidites may have come from a similar genus
Ischyosporites[120][121][116]
  • I. variegatus
  • I. bulbaceous
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the genus Dicksoniaceae inside Polypodiopsida. Tree fern spores
Example of extant Lophosoria specimens

Klukisporites[119][125][116]

  • K. pseudoreticulatus
  • K. variegatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Schizaeaceae inside Schizaeales.

Laevigatosporites[115]

  • L. laevis
  • L. sp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Blechnaceae inside Filicopsida.
Extant Sadleria, typical example of the Blechnaceae. Laevigatosporites spores are pretty similar to the extant ones of this genus
Maculatisporites[116]
  • M. granulatus
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
Spores Affinities with Cyatheaceae inside Cyatheales.

Maexisporites[121][125]

  • M. planatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Incertae Sedis affinities with the Pteridophyta.
Marattisporites[120][121][125]
  • M. scabratus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Marattiaceae inside Polypodiidae. Resemble the modern Marattia spores, probably belonging to a similar genus, related with large sized herbaceous ferns of humid environments.
Extant Marattia, typical example of the Marattiaceae. Marattisporites may have come from a similar genus
Matonisporites[120][121][125]
  • M. phlebopteroides
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Affinities with the Matoniaceae inside Polypodiidae. The so-called comb-ferns, found forming large colonies on humid settings.
Extant Matonia, typical example of the Matoniaceae. Matonisporites may have come from a similar genus
Thuringiasporites[120][121]
  • T. laevigatus
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Spores
Affinities with Cyatheaceae inside Cyatheales.

Todisporites[115]

  • T. hartzi
  • T. undans
  • T. goeppertianus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals

Spores

Affinities with the family Osmundaceae inside Polypodiopsida.

Trachytriletes[119][125]

  • T. dichotomus
  • T. filiciformis
  • T. lineatus
  • T. rugosus
  • Brody-Lubienia borehole.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores
Incertae Sedis affinities with the Pteridophyta. Uncertain Pteridophyte origin

Triancoraesporites[120][121]

  • T. communis
  • T. reticulatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Spores

Undulatisporites[116]

  • U. undulapolus
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Spores

Affinities with the Anemiaceae inside Schizaeales.

Extant Anemia caffrorum, typical example of the Anemiaceae. Undulatisporites spores are pretty similar to the extant ones of this genus

Cycadeoidophyta

[edit]
Genus Species Stratigraphic position Material Notes Images

Bennettistemon[120][121][115]

  • B. bursigerum
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with Williamsoniaceae the inside Bennettitales. Very abundant Pollen.

Reconstruction of Williamsonia, typical example of the Williamsoniaceae. Bennetistemon Pollen is considered to come from a similar genus

Chasmatosporites[117][118][125]

  • C. apertus
  • C. major
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals

Pollen

Affinities with the family Zamiaceae inside Cycadales. Is among the most abundant flora recovered on the upper section of the coeval Rya Formation, and was found to be similar to the pollen of the extant Encephalartos laevifolius.[126]

Extant Encephalartos laevifolius. Chasmatosporites maybe come from a related plant

Clavatipollenites[119][125][116]

  • C. hughesii
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the family Cycadaceae inside Cycadales. The structure of the exine of Clavatipollenites hughesii from Jurassic deposits is fundamentally different from that of Cretaceous grains referred to the same species, confirming observations made previously on the basis of analysis under the light microscope, and suggesting a possible derivation from cycadalean rather than angiospermous plants.[127]

Extant Cycas platyphylla. Clavatipollenites maybe come from a related plant

Cycadopites[116]

  • C. couperi
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Pollen

Cycadopollenites[116]

  • C.follicularis
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Pollen

Tricolpites[123][125]

  • T. troedssonii
  • T. gorzowiensis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Cycadeoidaceae inside Bennettitales.

Gnetophyta

[edit]
Genus Species Stratigraphic position Material Notes Images

Eucommiidites[125]

  • E. troedssonii
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Type Pollen of the Erdtmanithecales, that can be related with the Gnetales. Thick tectum, infratectum of small granules, indistinct or absent foot layer. Originally was thought to come from Angiosperms, latter reports suggest it come from arbustive Bennetites. It was recently found to come from Eucommiitheca, member of the enigmatic Erdtmanithecales, reinterpreted as an unusual gymnosperm grain with a single distal colpus flanked by two subsidiary lateral colps. Is very similar to the Pollen of the extant Ephedra and Welwitschia (mainly on the granular structure of the exine).[128]

Gnetophyta

[edit]
Genus Species Stratigraphic position Material Notes Images

Ginkgocycadophytus[116]

  • G. nitidus
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Pollen

Affinities with the family Karkeniaceae and Ginkgoaceae in Ginkgoales. Monosulcites was initially mistaken as Pollen coming from early Agiosperms, yet is likely from Ginkgoales

Extant Ginkgo, only surviving example of the Ginkgoaceae. Ginkgocycadophytus Pollen is pretty similar to the extant ones of this genus

Monosulcites[129]

  • M. carpentieri
  • M. minimus
  • M. spp.
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Brody-Lubienia Borehole

Pollen

Coniferophyta

[edit]
Genus Species Stratigraphic position Material Notes Images

Abietineaepollenites[129]

  • A. dunrobinensis
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Brody-Lubienia Borehole

Pollen

Affinities with the family Pinaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Extant Tsuga Cone, example of the Abietoideae. Pinaceae Pollen of the Ciechocinek fm is similar to the pollen found on this genus, hence the name

Agathoxylon[130]

  • A. agathiforme
  • “Elka” coal pit
  • “Kamilla” coal pit
  • “Zygmunt” coal pit
  • Isolated Logs
  • Isolated Branches
  • Isolated Indeterminate Woody Material
  • Coalified Fragments

Affinities with Hirmeriellaceae or Araucariaceae inside Pinales.

Agathoxylon

Applanopsipollenites[120][121][116]

  • A. dampieri
  • A. trilobatus
  • A. segmentatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Araucariaceae inside Coniferae. Resemble the pollen from the modern genus Agathis.

Extant Agathis Cone, example of the Araucariaceae. Applanopsipollenites is similar to the pollen found on this genus

Applanopsis[129]

  • A. trilobatus
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Pollen

Affinities with the Araucariaceae inside Coniferae. Resemble the pollen from the modern genus Agathis.

Araucariacites[117][118][125]

  • A. australis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals

Pollen

Affinities with the Araucariaceae inside Coniferae. Resemble the pollen from the modern genus Agathis.

Cerebropollenites[120][121][116]

  • C. macroverrucosus
  • C. serratus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with both Sciadopityaceae and Miroviaceae inside Pinopsida. This Pollen resemblance with extant Sciadopitys suggest that Miroviaceae can be an extinct lineage of sciadopityaceaous-like plants.[131]

Extant Sciadopitys. Cerebropollenites likely come from a related plant

Classopollis[120][121][116]

  • C. torosus
  • C. classoides
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Cheirolepidiaceae inside Coniferae. Is very abundant on hot and dry settings, found specially on the Brody-Lubienia Borehole.

Cupressacites[120][121][116]

  • C. minor
  • C. subgranulatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Cupressaceae inside Coniferae. The Pollen from this genus is similar to the present on the modern Fitzroya and Calocedrus.

Extant Calocedrus Cone, example of the Cupressaceae. Cupressacites is similar to the pollen found on this genus

Inaperturopollenites[120][121][125][116]

  • I. laevis
  • I. turbatus
  • I. elatoides
  • I. flavus
  • I. orbicularis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the family Pinaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Exesipollenites[120][121][116]

  • E. tumulus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Cupressaceae inside Coniferae. The Pollen from this genus is similar to the present on the modern Fitzroya and Calocedrus.

Ovalipollis[120][121][125]

  • O. breviformis
  • O. ovalis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen
Affinities with the family Pinaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Paleoconiferus[119][125]

  • P. asaccatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with Paleoconiferae inside Coniferae. Primitive Conifer and possible relict taxon

Patinasporites[119][125]

  • P. toralisides
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen
Affinities with the Patokaeaceae inside Coniferae. Pollen of the Conifer Patokaea silesiaca, know only from the Polish Upper Triassic (Norian) locality in Patoka.[132]

Perinopollenites[119][125][116]

  • P. elatoides
  • P. pseudosulcatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Cupressaceae inside Coniferae. The Pollen from this genus is similar to the present on the modern Fitzroya and Calocedrus.

Protoabies[115]

  • P. sp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen
Affinities with Paleoconiferae inside Coniferae. Primitive Conifer and possible relict taxon

Pseudopinus[115]

  • P. pectinella
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Pollen
Affinities with Paleoconiferae inside Coniferae. Primitive Conifer and possible relict taxon

Pityosporites[120][121][125]

  • P. haploxylon
  • P. minimus
  • P. alata
  • P. alatipollenites
  • P. silvestris
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen
Affinities with the family Pinaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Pristinuspollenites[117][118][125]

  • P. sulcatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals

Pollen

Affinities with the Podocarpaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Pseudopodocarpus[120][121]

  • P. sp.
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the Podocarpaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Quadraeculina[118][125]

  • Q. anellaeformis
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Blanowice Coals
  • Pollen

Affinities with the Podocarpaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Extant Podocarpus. Podocarpidites and Parvisaccites maybe come from a related plant

Rotundipollenites[119]

  • R. laevis
  • R. lineatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the family Pinaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

Sphaeripollenites[116]

  • S. subgranulatus
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Pollen

Affinities with the Cheirolepidiaceae inside Coniferae. Is very abundant on hot and dry settings, found specially on the Brody-Lubienia Borehole.

Zonalapollenites[119][125]

  • Z. aporosus
  • Z. acusus
  • Z. dampieri
  • Z. segmentatus
  • Z. trilobatus
  • Ciechocinek 58 Borehole
  • Gorzow Wielkopolski borehole
  • Mechowo borehole
  • Parkoszowice borehole
  • Cerpol-Kozlowice Borehole
  • Boroszów Borehole
  • Helenow Beds
  • Pollen

Affinities with the family Pinaceae inside Pinopsida. Conifer pollen from medium to large arboreal plants

References

[edit]
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