This article is about an Antarctica sedimentary sequence. For the geological feature, see
Mesa . For other uses, see
Meseta .
The La Meseta Formation is a sedimentary sequence deposited during much of the Paleogene on Seymour Island off the coast of the Antarctic Peninsula . It is noted for its fossils, which include both marine organisms and the only terrestrial vertebrate fossils from the Cenozoic of Antarctica.[ 2] [ 4]
In some treatments, the La Meseta Formation is restricted to just the older Thanetian to Lutetian -aged strata, with the younger Bartonian to Rupelian strata treated as the overlying Submeseta Formation .[ 5] [ 6] However, other papers instead treat the Submeseta Formation as an allomember of the La Meseta Formation.[ 7] [ 8]
La Meseta Formation lies unconformably on the Cretaceous Lopez de Bertodano Formation . It is an approximately 557 metres (1,827 ft) thick sequence of poorly consolidated sandstones and siltstones . The depositional environment was probably coastal, deltaic or estuarine in character. The top of the sequence is an erosional unconformity to Pleistocene glacial gravels.[ 3] [ 2] La Meseta Formation is one of the sequences that make up the fill of the Late Jurassic to Paleogene James Ross Basin .[ 3]
The terrestrial environment surrounding the deposition area is thought to have been a temperate polar forest, including podocarp and araucarian conifers, as well as Nothofagus .[ 9] [ 10] Most of the fossilized woods and flowers discovered on Seymour Islands consist of extinct species of conifer trees and lilies during warm climate.[ 11] [ 12]
La Meseta Formation is extremely rich in fossils. Among mammals, the meridiungulata Antarctodon and Trigonostylops have been found in the formation.[ 13] [ 14] as well as marsupial Derorhynchidae , Microbiotheria , and polydolopimorphia .[ 15] [ 16] It is famous for its penguin fossils, for example the two genera Archaeospheniscus and Palaeeudyptes .[ 17] [ 18] Other bird fossils include Dasornis , a genus of pseudotooth birds . There is also an abundance of trace fossils. Diplocraterion , Helminthopsis , Muensteria , Oichnus , Ophiomorpha , Skolithos , Teredolites and Zapfella have been described.[ 19] Over 35 species and 26 families of fish, which includes sharks, have been described from the Ypresian Cucullaea bed.[ 3] [ 20]
Color key
Notes Uncertain or tentative taxa are in small text ; crossed out taxa are discredited.
Astrapotheria
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Antarctodon
A. sobrali
Locality DVP 2/84 and IAA 1/90.
Cucullaea I allomember (Telm 5).
MLP 08-XI-30-1, an isolated right p4 or m1.
An astrapothere
Trigonostylops
T. sp.
Cucullaea I allomember (Telm 5).
An astrapothere
Litopterna
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Notiolofos
N. arquinotiensis
Locality IAA 1/13 and DPV 16/84.
Acantilados II, Campamento, Cucullaea I Allomember (Telm 4) and Submeseta Member (Telm 7).
A Brachyodont lower right molar fragment, probably m1 or m2, preserves a portion of the talonid with most of the lingual side (MLP 13-I-25-1) and left M3 incomplete (MLP 95-I-10-6).
A sparnotheriodontid litoptern .
N. regueroi
Locality IAA 2/16
Lower coquina bed of Cucullaea I Allomember.
IAA-PV 173, Isolated complete left m3.
Victorlemoinea
V. sp.
Cucullaea I allomember (Telm 5).
A sparnotheriodontid litoptern.
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Basilosauridae [ 21]
Indeterminate
Llanocetus
L. denticrenatus
Locality DPV 10/84 and DVP 13/24.
Unit III Member and Telm 7.
A partial mandible with two teeth (specimen USNM 183022) and an endocast of the braincase.
A toothed baleen whale.
Derorhynchidae
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Derorhynchus [ 22]
D. minutus [ 22]
Locality IAA 1/90.
Allomember Cucullaea I (Telm 5).
MLP 96-1-5-44, incomplete right horizontal ramus of the dentary fragment with m2-3.
Pauladelphys [ 22] [ 23]
P. juanjoi [ 22] [ 23]
Locality IAA 2/95 and IAA 1/90.
Allomember Cucullaea I (Telm 5).
An isolated lower molar (MLP 95-1-10-2) and an upper left molar (MLP 96-1-5-44).
Xenostylos
X. peninsularis [ 22]
Locality IAA 1/90.
Allomember Cucullaea I (Telm 5).
MLP 94-111-15-10, an isolated upper right molar.
Microbiotheria
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Marambiotherium [ 22]
M. glacialis [ 22]
Locality IAA 1/90 and RV-8200.
Cucullaea I allomember (Telm 5).
MLP 95-1-10-1, a fragment of a right mandibular ramus with a complete m4 and the posterior alveolus of m3 and MLP 88-1-1-1, an edentulous left mandibular ramus with the alveoli for p3-m4.
A microbiothere , related to the modern monito del monte .
Woodburnodon
W. casei
Locality IAA 1/95.
Cucullaea I allomember (Telm 5).
MLP 04-III-1-2, an isolated, worn upper right molar (M2 or M3).
A microbiothere, related to the modern monito del monte.
Polydolopimorphia
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Antarctodolops
A. dailyi
Locality IAA 2/95, DPV 2/84, and DPV 6/84.
Cucullaea I allomember (Telm 5).
UCR 20910, a left dentary with p3-m2.
A polydolopimorphian metatherian.
A. mesetaense
Locality IAA 1/90 and DPV 6/84.
Cucullaea I allomember (Telm 5).
Right dentary fragments.
Perrodelphys
P. coquinense [ 22]
Locality IAA 1/90.
Cucullaea I allomember (Telm 5).
MLP 96-1-5-11, an isolated left lower molar.
A polydolopimorphian metatherian.
Polydolops
P. dailyi
Locality IAA 1/90 & DPV 3/84.
Cucullaea I allomember (Telm 5).
A polydolopimorphian metatherian.
P. seymouriensis
Locality IAA 1/90 & DPV 3/84.
Cucullaea I allomember (Telm 5).
P. thomasi
Locality IAA 1/90 & DPV 3/84.
Cucullaea I allomember (Telm 5).
Pujatodon
P. ektopos [ 24]
Locality IAA 1/90.
Cucullaea I Allomember (Telm 5).
Specimen MLP 14-I-10-20, a lower left molar (m2 or m3) partially worn.
A polydolopimorphian metatherian.
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Sudamerica
S. ameghinoi
Locality IAA 1/90 upper level.
Cucullaea I Allomember.
MLP 95-I-10-5, anterior part of a left dentary with the rodent-like incisor partially preserved.
A gondwanathere
Meridiolestida ?[ 25]
Indeterminate
Locality IAA 1/90 upper level.
Cucullaea I Allomember.
A single tooth, now lost.[ 25]
Xenarthra ?
Indeterminate.
Locality S124.
Cucullaea I Allomember/ Telm 4.
TMM 44190-1, left metacarpal II and A phalanx and a partial tooth[ 25]
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Anthropornis
A. grandis
Locality IAA 4/12, IAA 1/90, DPV 13/84, and DVP 2/84.
Telm 4 member, Telm 7 member, and Submeseta Allomember.
Humerus: MLP 93-X-1-4 (proximal epiphysis), MLP 82-IV-23-4 (proximal epiphysis), MLP 83-I-1-190(proximal epiphysis) and MLP 88-I-1-463(proximal epiphysis).
A giant penguin .
Anthropornis Palaeeudyptes
A. nordenskjoldi [ 23]
Submeseta Allomember.
Humerus: MLP CX-60-25 (proximal epiphysis), MLP 83-V-30-5 (diaphysis) and MLP 93-X-1-104(complete humerus).
Archaeospheniscus
A. lopdelli
Submeseta Allomember.
A giant penguin.
A. wimani
Submeseta Allomember.
Delphinornis [ 23]
D. arctowskii
DPV 14/84
Cucullaea I Allomember (Telm 7).
A penguin.
D. graclis
DPV 14/84
Cucullaea I Allomember (Telm 7).
D. larseni
DPV 13/84 and DPV 14/84.
Cucullaea I Allomember (Telm 5 and Telm 7) and Submeseta Allomember.
Marambiornis
M. exilis
DVP 2/84, IAA 1/12, and ZPAL 4
Telm 7 Member.
A limb element (right tarsometatarsus ).
A penguin.
Mesetaornis
M. polaris
Telm 7 member.
A nearly complete right femur and two distal left tibiotarsi.
A penguin.
Orthopteryx
O. gigas [ 18]
A penguin.
Palaeeudyptes
P. antarcticus
Cucullaea I Allomember (Telm 3, Telm 4, Telm 5, and Telm 7) and Submeseta Allomember.
A giant penguin.
P. gunnari [ 23]
DVP 2/84, DVP 10/84, DVP 13/84, DVP 14/84, DVP 15/84, and ZPAL 4.
Cucullaea I Allomember (Telm 3, Telm 5, and Telm 7) and Submeseta Allomember.
Multiple specimens consist of coracoids, ulna, and humerus.
P. klekowskii
DVP 2/84, DVP 10/84, DVP 13/84, DVP 14/84, DVP 16/84, and ZPAL 4.
Cucullaea I Allomember (Telm 3, Telm 5, and Telm 7) and Submeseta Allomember.
Several specimens consist of humerus, coracoid, ulna, and tibiotarsus.
Wimanornis
W. seymourensis
DPV 14/84.
Cucullaea I Allomember (Telm 7) and Submeseta Allomember.
A penguin.
Tonniornis
T. mesetaensis
DPV 14/84.
Cucullaea I Allomember (Telm 7) and Submeseta Allomember.
A penguin.
T. minimum
DPV 14/84.
Cucullaea I Allomember (Telm 7) and Submeseta Allomember.
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Calyptocephalella
C. sp. [ 33]
Locality IAA 2/95.
Cucullaea I Allomember (Telm 5).
The fragmentary right ilium (NRM-PZ B282) and a skull bone (NRM-PZ B281).
A calyptocephalellid frog
Taxa
Species
Locality
Stratigraphic position
Material
Notes
Images
Antarcticeras
A. nordenskjoeldi
NRM 8, below IAA 1/90 (Ungulate Site).
Cucullaea I shell bed, Telm 4.
A fragmentary preserved straight shell.
An enigmatic cephalopod that is either considered a descendant of the orthocerids or an oegopsid squid.
Euciphoceras
E . sp.
NRM 8, below IAA 1/90 (Ungulate Site).
Cucullaea I shell bed, Telm 4.
A fragmentary preserved straight shell.
A nautiloid .
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