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Wild horse

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This article is about the species Equus ferus. For free-roaming horses descended from domesticated ancestry, see Feral horse. For other uses, see Wild horse (disambiguation).

Wild horse
Temporal range: earliest Middle Pleistocene-Recent 0.8–0 Ma
↓
Top left: Equus ferus caballus (horses)
Top right: Equus ferus przewalskii (Przewalski's horse)

Below left: Equus ferus ferus† (tarpan)
Below right: Equus ferus fossil from 9100 BC

Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Genus: Equus
Subgenus: Equus
Species:
E. ferus
Binomial name
Equus ferus
Boddaert, 1785
Subspecies

The wild horse (Equus ferus) is a species of the genus Equus, which includes as subspecies the modern domesticated horse (Equus ferus caballus) as well as the endangered Przewalski's horse (Equus ferus przewalskii, sometimes treated as a separate species i.e. Equus przewalskii).[2][3] The European wild horse, also known as the tarpan, that went extinct in the late 19th or early 20th century has previously been treated as the nominate subspecies of wild horse, Equus ferus ferus, but more recent studies have cast doubt on whether tarpans were truly wild or if they actually were feral horses or hybrids.[4][5][6]

Other subspecies of Equus ferus may have existed and could have been the stock from which domesticated horses are descended.[7] Przewalski's horse had reached the brink of extinction, but was reintroduced successfully into the wild.[8] The tarpan became extinct in the 19th century, but is theorized to have been present on the steppes of Eurasia at the time of domestication.[9][10][11][12] Since the extinction of the tarpan, attempts have been made to reconstruct its phenotype using domestic horses, resulting in horse breeds such as the Heck horse.[13][14] However, the genetic makeup and foundation bloodstock of those breeds is substantially derived from domesticated horses, so these breeds possess domesticated traits.

The term "wild horse" is also used colloquially in reference to free-roaming herds of feral horses; for example, the mustang in the United States,[15] and the brumby in Australia.[16] These feral horses are untamed members of the domestic horse (Equus caballus), not to be confused with the truly "wild" horse subspecies extant into modern times.

Distribution

[edit]

Wild horses were formerly widespread across the Old World during the Late Pleistocene and early Holocene, occurring from Western Europe and North Africa (where remains are referred to as Equus algericus), to the far north of Siberia (where they are referred to as Equus lenensis) to East Asia.[17] Today the only living wild horse subspecies, Przewalski's horse, which was formerly extinct in the wild, has been reintroduced to small areas spanning from its former distribution in northeast China, Mongolia[18] and Kazakhstan,[19] as well as areas outside of its historical distribution, including Spain.[20]

Description

[edit]
See also: Horse § Anatomy

Across time and space, wild horses show considerable variability in body size and limb dimensions, likely as adaptations to local environmental and climatic conditions.[21][22] The living Przewalski's horse has an estimated body mass of 250–360 kilograms (550–790 lb),[23] though the body mass of Przewalski's horse varies considerably according to rythmic annual cycles.[24] Some extinct wild horse populations are thought to have been considerably larger, with some Pleistocene European wild horses (including those assigned to Equus (ferus) mosbachensis and Equus (ferus) latipes) suggested to have body masses of around 500–607 kilograms (1,102–1,338 lb).[25][26][22]ST2 Earlier Middle Pleistocene horses (such as those assigned to E. (ferus) mosbachensis) differ from later ones in some aspects of the anatomy of the limbs and teeth.[27]

Ecology

[edit]

Wild horses show considerable environmental tolerances, having historically inhabited environments ranging from temperate forest to steppe[27] (including the prehistoric mammoth steppe biome[28]) and tundra,[29] though in general they tend to show a preference for open environments.[30] Horses are typically considered to be grazers.[30] They may have seasonal food preferences, as seen in the Przewalski's horse,[31] which historically consumed browse like shrubs during the winter months due to being forced into suboptimal habitat by human pressure.[32] Some extinct Pleistocene wild horse populations that inhabited forested environments show dental wear suggesting them to have been mixed feeders or even predominantly browsers during certain times of the year, though this may be reflecting the consumption of low growing forbs rather than shrubs.[22][33][34]

Wild horses live in herds with a social hierarchy,[35] formed by a dominant adult male or sometimes multiple males (harem stallions), as well as several mares and their offspring. The harem stallion aggressively defends his herd/harem against rival males.[36] Upon reaching adulthood, both male and female horses disperse to other herds to avoid inbreeding,[35] with young adult males also forming batchelor groups[36] when they are around 3 years of age. In batchelor groups male horses engage in play and ritual behaviour, with the group forming a hierarchy.[37] Mare Przewalski's horses tend to begin giving birth around 3 years of age,[38] with a gestation period of around 12 months, primarily giving birth in the spring or summer.[39] Przewalski's stallions generally leave batchelor groups to begin breeding at around 5-6 years of age.[37]

In modern times the main predator of wild horses in Eurasia are wolves,[40] though during the Pleistocene they had other predators such as cave hyenas.[41]

Subspecies and their history

[edit]

E. ferus has had several subspecies, those of which survived into modern times are:[42]

  • The domesticated horse (Equus ferus caballus).
  • The Eurasian wild horse (Equus ferus ferus), incorrectly listed as Equus caballus ferus in MSW 3; originally considered synonymous with the tarpan, though recent research has cast doubt on this. Horses identified as tarpans were found in Europe and western Asia before the last surviving animals —possibly hybrids by that time — became effectively extinct in the late 19th century. The last specimen died in 1909 whilst in captivity in an estate in Poltava Governorate, Russian Empire.
  • Przewalski's horse (Equus ferus przewalskii), incorrectly listed as Equus caballus przewalskii in MSW 3; also known as the Mongolian wild horse or takhi, it is native to Central Asia and the Gobi Desert. It is sometimes considered its own species, Equus przewalskii.

The latter two are the only never-domesticated "wild" groups that survived into historic times. However, other subspecies of Equus ferus may have existed.[7]

In the Late Pleistocene epoch, there were several other subspecies of E. ferus which have all since gone extinct. The exact categorization of Equus remains into species or subspecies is a complex matter and the subject of ongoing work.[27]

Evolutionary history and taxonomy

[edit]
Equus ferus fossil from 9100 BC found near Odense, at the Zoological Museum in Copenhagen
Main article: Evolution of the horse

The horse family Equidae evolved in North America, with the genus Equus appearing on the continent during the Pliocene (5.3-2.6 million years ago).[43] Horses are thought to have diverged from the ancestors of zebras and asses around 4 million years ago.[44] Around 900-800,000 years ago, at the Early-Middle Pleistocene boundary, the ancestors of Eurasian wild horses crossed over the Bering Land Bridge from North America.[45] Early horses in Eurasia are referred to species like Equus mosbachensis.[27] North American caballine horses (which are variously referred to species like Equus scotti and Equus lambei though the true number of species is uncertain) which genetic evidence has confirmed are closely related to Eurasian horses (with some authors treating all North American horses as part of E. ferus) would persist on the continent until they became extinct as part of the end-Pleistocene extinction event along with most other large mammals in the Americas (including other equines like Haringtonhippus and Hippidion) around 12,000 years ago. Ancient DNA analysis suggests North American horses were largely genetically distinct from Eurasian horses, with limited and intermittent interbreeding between the two populations following the initial dispersal across the Bering Land Bridge.[45] Equus neogeus from the Pleistocene of South America may also represent a lineage of true caballine horses.[46]

Currently, three subspecies that lived during recorded human history are recognized.[42] One subspecies is the widespread domestic horse (Equus ferus caballus),[42] as well as two wild subspecies: the recently extinct European wild horse (E. f. ferus) and the endangered Przewalski's horse (E. f. przewalskii).[10][11][42] A 2015 study determined that the Przewalski and domesticated horse lineages diverged from a common ancestor about 45,000 years ago.[47]

Genetically, the pre-domestication horse, E. ferus, and the domesticated horse, E. caballus, form a single homogeneous group (clade) and are genetically indistinguishable from each other.[48][49][50][51] The genetic variation within this clade shows only a limited regional variation, with the notable exception of Przewalski's horse.[48][49][50][51] Przewalski's horse has several unique genetic differences that distinguish it from the other subspecies, including 66 instead of 64 chromosomes,[10][52] unique Y-chromosome gene haplotypes,[53] and unique mtDNA haplotypes.[54][55]

Besides genetic differences, osteological evidence from across the Eurasian wild horse range, based on cranial and metacarpal differences, indicates the presence of only two subspecies in postglacial times, the tarpan and Przewalski's horse.[7][56] A study in 2011 of DNA from bones of pre-domestication horses found that all were either bay, black or leopard-spotted.[57]

Scientific naming of the species

[edit]

In some sources including MSW 3 (2005), the domesticated and wild horses were considered a single species, with the valid scientific name for such a single horse species being Equus ferus,[58] although MSW erroneously used E. caballus for this (enlarged) taxon on account of a mis-interpretation of the then-recent ICZN ruling on the matter,[59] refer Groves & Grubb, 2011.[60] The wild tarpan subspecies is E. f. ferus, Przewalski's horse is E. f. przewalskii, while the domesticated horse is nowadays normally (but not exclusively) treated as a separate species E. caballus. The rules for the scientific naming of animal species are determined in the International Code of Zoological Nomenclature, which stipulates that the oldest available valid scientific name is used to name the species.[61] Previously, when taxonomists considered domesticated and wild horse two subspecies of the same species, the valid scientific name was Equus caballus Linnaeus 1758,[62] with the subspecies labeled E. c. caballus (domesticated horse), E. c. ferus Boddaert, 1785 (tarpan) and E. c. przewalskii Poliakov, 1881 (Przewalski's horse).[63] However, in 2003, the International Commission on Zoological Nomenclature decided that the scientific names of the wild species have priority over the scientific names of domesticated species, therefore mandating the use of Equus ferus for both the wild and the domesticated horse if the two taxa are considered conspecific.[64]

Przewalski's horse

[edit]
Przewalski's horse in Hungary

Przewalski's horse occupied the eastern Eurasian Steppes, perhaps from the Ural Mountains to Mongolia, although the ancient border between tarpan and Przewalski's distributions has not been clearly defined.[65] Przewalski's horse was limited to Dzungaria and western Mongolia in the same period, and became extinct in the wild during the 1960s, but was reintroduced in the late 1980s to two preserves in Mongolia.[66] Although earlier researchers such as Marija Gimbutas theorized that the horses of the Chalcolithic period were Przewalski's, a 2003 study indicated that the Przewalski's horse is not an ancestor to modern domesticated horses.[67][68]

In 2018, a DNA study revealed that the horses raised for meat and milk[69] by the Botai culture 5500 years ago were Przewalski's horses. The paper claims specifically that modern Przewalski's horses are the feral descendents of the domesticated[70][71] Botai horse, although it is also possible both groups could have descended separately from the same ancient wild Przewalski's horses.[72][73]

Przewalski's horse is still found today, though it is an endangered species and for a time was considered extinct in the wild.[55] Roughly 2000 Przewalski's horses are in zoos around the world.[74] A small breeding population has been reintroduced in Mongolia.[75][76] As of 2005, a cooperative venture between the Zoological Society of London and Mongolian scientists has resulted in a population of 248 animals in the wild.[77]

Przewalski's horse has some biological differences from the domestic horse; unlike domesticated horses and the tarpan, which both have 64 chromosomes, Przewalski's horse has 66 chromosomes due to a Robertsonian translocation.[78] However, the offspring of Przewalski and domestic horses are fertile, possessing 65 chromosomes.[79]

Feral horses

[edit]
Main articles: Feral horse, semi-feral, and feral
See also: List of megafauna discovered in modern times § Status unclear
Semi-feral Exmoor ponies. Though popularly called "wild" horses, feral and semi-feral horses had ancestors that were domesticated.

Horses that live in an untamed state but have ancestors that have been domesticated are called "feral horses".[80] For instance, when the Spanish reintroduced the horse to the Americas, beginning in the late 15th century,[81] some horses escaped, forming feral herds; the best-known being the mustang.[82] Similarly, the brumby descended from horses strayed or let loose in Australia by English settlers.[83] Isolated populations of feral horses occur in a number of places, including Bosnia, Croatia, New Zealand, Portugal, Scotland and a number of barrier islands along the Atlantic coast of North America from Sable Island off Nova Scotia, to Cumberland Island, off the coast of Georgia.[84] Even though these are often referred to as "wild" horses, they are not truly "wild" if wildness is defined as having no domesticated ancestors.[80]

In 1995, British and French explorers encountered a new population of horses in the Riwoche Valley of Tibet, unknown to the rest of the world, but apparently used by the local Khamba people.[85] It was speculated that the Riwoche horse might be a relict population of wild horses,[86] but testing did not reveal genetic differences with domesticated horses,[87] which is in line with news reports indicating that they are used as pack and riding animals by the local villagers.[88] These horses only stand 12 hands (48 inches, 122 cm) tall and are said to resemble the images known as "horse no 2" depicted in cave paintings alongside images of Przewalski's horse.[87]

Relationship with humans

[edit]

Archaic humans hunted horses hundreds of thousands of years before the dispersal of modern humans across Eurasia during the Last Glacial Period. Examples of sites demonstrating horse butchery by archaic humans include the Boxgrove site in southern England dating to around 500,000 years ago, where horse bones with cut marks (with a horse scapula possibly exhibiting a spear wound[89]) are associated with Acheulean stone tools made by Homo heidelbergensis,[90][91] the Schöningen site in Germany (also thought to have been created by Homo heidelbergensis) dating to around 300,000 years ago, where butchered horses are associated with wooden spears (the Schöningen spears, amongst the oldest known wooden spears),[92][93] as well as the Lingjing site in Henan, China dating to 125-90,000 years ago.[94] During the Upper Palaeolithic, there is evidence for the hunting of horses by modern humans in Europe,[95] as well as Asia.[96] Early Paleoindians in North America hunted the continent's native horses shortly prior to their extinction.[97] During the 3rd millennium BC, horses were domesticated on the western Eurasian steppes, with domestic horses spreading across Eurasia around 2000 BC.[98]

See also

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References

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Citations

[edit]
  1. ^ King, S.R.B.; Boyd, L.; Zimmermann, W. & Kendall, B.E. (2016) [errata version of 2015 assessment]. "Equus ferus". IUCN Red List of Threatened Species. 2015: e.T41763A97204950. doi:10.2305/IUCN.UK.2015-2.RLTS.T41763A45172856.en. Retrieved 17 February 2022.
  2. ^ Grubb, P. (2005). "Order Perissodactyla". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 630–631. ISBN 978-0-8018-8221-0. OCLC 62265494.
  3. ^ "Explore the Database". www.mammaldiversity.org. Retrieved 20 August 2021.
  4. ^ Tadeusz Jezierski, Zbigniew Jaworski: Das Polnische Konik. Die Neue Brehm-Bücherei Bd. 658, Westarp Wissenschaften, Hohenwarsleben 2008, ISBN 3-89432-913-0
  5. ^ "Tarpan". Britannica. Retrieved 7 March 2023.
  6. ^ "The Przewalskii and Tarpan Horses". Retrieved 7 March 2023.
  7. ^ a b c Colin Groves, 1986, "The taxonomy, distribution, and adaptations of recent Equids", In Richard H. Meadow and Hans-Peter Uerpmann, eds., Equids in the Ancient World, volume I, pp. 11-65, Wiesbaden: Ludwig Reichert Verlag.
  8. ^ Goldman, Jason G. "10 Things You Didn't Know About Przewalski's Horses". Scientific American Blog Network.
  9. ^ "The First Horses: The Przewalskii and Tarpan Horses", The legacy of the horse, International Museum of the Horse, archived from the original on 30 October 2007, retrieved 18 February 2009
  10. ^ a b c Groves, Colin P. (1994). Boyd, Lee and Katherine A. Houpt (ed.). The Przewalski Horse: Morphology, Habitat and Taxonomy. Vol. Przewalski's Horse: The History and Biology of an Endangered Species. Albany, New YorkColin P. Groves: State University of New York Press.
  11. ^ a b Kavar, Tatjana; Peter Dovč (2008). "Domestication of the horse: Genetic relationships between domestic and wild horses". Livestock Science. 116 (1–3): 1–14. doi:10.1016/j.livsci.2008.03.002.
  12. ^ Bowling, Ann T.; Anatoly Ruvinsky (2000). "Genetic Aspects of Domestication, Breeds and Their Origin". In Ann T. Bowling; Anatoly Ruvinsky (eds.). The Genetics of the Horse. CABI Publishing. ISBN 978-0-85199-429-1.
  13. ^ "Tarpan or Heck Horse". horsehints.org.
  14. ^ "Rare horse breed proves crucial to delicate ecosystem - Features". Horsetalk.co.nz. 1 June 2012. Archived from the original on 9 November 2020. Retrieved 28 May 2017.
  15. ^ "Black Hills Wild Horse Sanctuary". Black Hills Wild Horse Sanctuary.
  16. ^ "The Brumbies – Australian Wild Horses › Wild Horses and Mustangs .com". www.wildhorsesandmustangs.com.
  17. ^ Cirilli, Omar; Machado, Helena; Arroyo-Cabrales, Joaquin; et al. (24 August 2022). "Evolution of the Family Equidae, Subfamily Equinae, in North, Central and South America, Eurasia and Africa during the Plio-Pleistocene". Biology. 11 (9): 1258. doi:10.3390/biology11091258. ISSN 2079-7737. PMC 9495906. PMID 36138737.
  18. ^ Ji, Shengnan; Zhu, Yanpeng; Cui, Shaopeng; et al. (September 2022). "The road home for Przewalski's horse in China". Oryx. 56 (5): 652. doi:10.1017/S0030605322000758. ISSN 0030-6053.
  19. ^ "Przewalski's horses return to central Kazakhstan after nearly 200-year absence". Frankfurt Zoological Society. 6 June 2024.
  20. ^ "La renaturalización trae al Alto Tajo los únicos caballos Przewalski que pastarán en libertad en Europa Occidental". Rewilding Spain. 30 November 2023. Retrieved 5 May 2024.
  21. ^ van Asperen, Eline Naomi (November 2010). "Ecomorphological adaptations to climate and substrate in late Middle Pleistocene caballoid horses". Palaeogeography, Palaeoclimatology, Palaeoecology. 297 (3–4): 584–596. Bibcode:2010PPP...297..584V. doi:10.1016/j.palaeo.2010.09.007.
  22. ^ a b c Saarinen, Juha; Cirilli, Omar; Strani, Flavia; et al. (26 February 2021). "Testing Equid Body Mass Estimate Equations on Modern Zebras—With Implications to Understanding the Relationship of Body Size, Diet, and Habitats of Equus in the Pleistocene of Europe". Frontiers in Ecology and Evolution. 9. doi:10.3389/fevo.2021.622412. hdl:11573/1505920. ISSN 2296-701X.
  23. ^ "Przewalski's horse". Smithsonian's National Zoo and Conservation Biology Institute. Retrieved 9 December 2024.
  24. ^ Scheibe, Klaus M.; Streich, Wolf J. (October 2003). "Annual Rhythm of Body Weight in Przewalski Horses (Equus ferus przewalskii)". Biological Rhythm Research. 34 (4): 383–395. Bibcode:2003BioRR..34..383S. doi:10.1076/brhm.34.4.383.26227. ISSN 0929-1016.
  25. ^ Pushkina, Diana (19 November 2024). "Paleoecology and body mass distribution of horses in northern Eurasia during the Pleistocene". Annales Zoologici Fennici. 61 (1). doi:10.5735/086.061.0118. ISSN 0003-455X.
  26. ^ Nacarino-Meneses, Carmen; Orlandi-Oliveras, Guillem (4 May 2021). "The life history of European Middle Pleistocene equids: first insights from bone histology". Historical Biology. 33 (5): 672–682. Bibcode:2021HBio...33..672N. doi:10.1080/08912963.2019.1655011. ISSN 0891-2963.
  27. ^ a b c d Boulbes, Nicolas; van Asperen, Eline N. (2019). "Biostratigraphy and Palaeoecology of European Equus". Frontiers in Ecology and Evolution. 7: 301. doi:10.3389/fevo.2019.00301. ISSN 2296-701X.
  28. ^ Schwartz-Narbonne, R.; Longstaffe, F.J.; Kardynal, K.J.; et al. (July 2019). "Reframing the mammoth steppe: Insights from analysis of isotopic niches". Quaternary Science Reviews. 215: 1–21. Bibcode:2019QSRv..215....1S. doi:10.1016/j.quascirev.2019.04.025.
  29. ^ Boeskorov, Gennady G.; Potapova, Olga R.; Protopopov, Albert V.; et al. (July 2018). "A study of a frozen mummy of a wild horse from the Holocene of Yakutia, east Siberia, Russia". Mammal Research. 63 (3): 307–314. doi:10.1007/s13364-018-0362-4. ISSN 2199-241X.
  30. ^ a b Sommer, Robert S.; Benecke, Norbert; Lõugas, Lembi; et al. (November 2011). "Holocene survival of the wild horse in Europe: a matter of open landscape?". Journal of Quaternary Science. 26 (8): 805–812. Bibcode:2011JQS....26..805S. doi:10.1002/jqs.1509. ISSN 0267-8179.
  31. ^ Kateryna Slivinska; Grzegorz Kopij (2011). "Diet of the Przewalski's horse Equus przewalskii in the Chernobyl Exclusion Zone" (PDF). Polish Journal of Ecology. 59 (4): 841–847. Archived from the original (PDF) on 29 October 2013. Retrieved 29 October 2013.
  32. ^ Kaczensky, Petra; Burnik Šturm, Martina; Sablin, Mikhail V.; et al. (20 July 2017). "Stable isotopes reveal diet shift from pre-extinction to reintroduced Przewalski's horses". Scientific Reports. 7 (1): 5950. Bibcode:2017NatSR...7.5950K. doi:10.1038/s41598-017-05329-6. ISSN 2045-2322. PMC 5519547. PMID 28729625.
  33. ^ Rivals, Florent; Julien, Marie-Anne; Kuitems, Margot; et al. (December 2015). "Investigation of equid paleodiet from Schöningen 13 II-4 through dental wear and isotopic analyses: Archaeological implications". Journal of Human Evolution. 89: 129–137. Bibcode:2015JHumE..89..129R. doi:10.1016/j.jhevol.2014.04.002. PMID 25242064.
  34. ^ Rivals, Florent; Lister, Adrian M. (August 2016). "Dietary flexibility and niche partitioning of large herbivores through the Pleistocene of Britain". Quaternary Science Reviews. 146: 116–133. Bibcode:2016QSRv..146..116R. doi:10.1016/j.quascirev.2016.06.007.
  35. ^ a b Tatin, L.; King, S. R. B.; Munkhtuya, B.; et al. (February 2009). "Demography of a socially natural herd of Przewalski's horses: an example of a small, closed population". Journal of Zoology. 277 (2): 134–140. doi:10.1111/j.1469-7998.2008.00520.x. ISSN 0952-8369.
  36. ^ a b Bernátková, Anna; Ceacero, Francisco; Oyunsaikhan, Ganbaatar; et al. (June 2023). "Novel observation of play behaviour between a harem holder and a bachelor group of Przewalski's horses in the wild". Acta Ethologica. 26 (2): 145–150. doi:10.1007/s10211-023-00421-9. ISSN 0873-9749.
  37. ^ a b Bourjade, M.; Tatin, L.; King, S.R.B.; Feh, C. (January 2009). "Early reproductive success, preceding bachelor ranks and their behavioural correlates in young Przewalski's stallions". Ethology Ecology & Evolution. 21 (1): 1–14. Bibcode:2009EtEcE..21....1B. doi:10.1080/08927014.2009.9522507. ISSN 0394-9370.
  38. ^ Rödel, Heiko G.; Ibler, Benjamin; Ozogány, Katalin; Kerekes, Viola (December 2023). "Age-specific effects of density and weather on body condition and birth rates in a large herbivore, the Przewalski's horse". Oecologia. 203 (3–4): 435–451. Bibcode:2023Oecol.203..435R. doi:10.1007/s00442-023-05477-9. ISSN 0029-8549. PMC 10684615. PMID 37971561.
  39. ^ Chen, Jinliang; Weng, Qiang; Chao, Jie; et al. (2008). "Reproduction and Development of the Released Przewalski's Horses (Equus przewalskii) in Xinjiang, China". Journal of Equine Science. 19 (1): 1–7. doi:10.1294/jes.19.1. ISSN 1340-3516. PMC 4019202. PMID 24833949.
  40. ^ Usukhjargal, Dorj; Henkens, RenéJ H. G.; Boer, Willem F. de; et al. (28 August 2009). "Wolf Predation Among Reintroduced Przewalski Horses in Hustai National Park, Mongolia". Journal of Wildlife Management. 73 (6): 836–843. doi:10.2193/2008-027. S2CID 85877321.
  41. ^ Diedrich, Cajus G. (15 June 2010). "Specialized horse killers in Europe: Foetal horse remains in the Late Pleistocene Srbsko Chlum-Komín Cave hyena den in the Bohemian Karst (Czech Republic) and actualistic comparisons to modern African spotted hyenas as zebra hunters". Quaternary International. 220 (1–2): 174–187. Bibcode:2010QuInt.220..174D. doi:10.1016/j.quaint.2010.01.023.
  42. ^ a b c d Don E. Wilson; DeeAnn M. Reeder, eds. (2005). "Equus caballus". Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. Retrieved 12 February 2009.
  43. ^ Beever, Erik A. (2022). "Equus caballus [ISC] (horse)". Cabi Compendium. CABI Compendium. doi:10.1079/cabicompendium.119345.
  44. ^ Cai, Dawei; Zhu, Siqi; Gong, Mian; et al. (11 May 2022). "Radiocarbon and genomic evidence for the survival of Equus Sussemionus until the late Holocene". eLife. 11. doi:10.7554/eLife.73346. ISSN 2050-084X. PMC 9142152. PMID 35543411.
  45. ^ a b Vershinina, Alisa O.; Heintzman, Peter D.; Froese, Duane G.; et al. (December 2021). "Ancient horse genomes reveal the timing and extent of dispersals across the Bering Land Bridge". Molecular Ecology. 30 (23): 6144–6161. Bibcode:2021MolEc..30.6144V. doi:10.1111/mec.15977. hdl:10037/24463. ISSN 0962-1083. PMID 33971056.
  46. ^ Barrón-Ortiz, Christina I.; Avilla, Leonardo S.; Jass, Christopher N.; et al. (12 September 2019). "What Is Equus? Reconciling Taxonomy and Phylogenetic Analyses". Frontiers in Ecology and Evolution. 7: 343. doi:10.3389/fevo.2019.00343. ISSN 2296-701X.
  47. ^ Der Sarkissian, Clio; Ermini, Luca; Schubert, Mikkel; et al. (October 2015). "Evolutionary Genomics and Conservation of the Endangered Przewalski's Horse". Current Biology. 25 (19): 2577–2583. Bibcode:2015CBio...25.2577D. doi:10.1016/j.cub.2015.08.032. hdl:10230/33065. PMC 5104162. PMID 26412128.
  48. ^ a b Weinstock, J.; et al. (2005). "Evolution, systematics, and phylogeography of Pleistocene horses in the New World: a molecular perspective". PLOS Biology. 3 (8): e241. doi:10.1371/journal.pbio.0030241. PMC 1159165. PMID 15974804.
  49. ^ a b Orlando, Ludovic; Male, Dean; Alberdi, Maria Teresa; et al. (9 April 2008). "Ancient DNA Clarifies the Evolutionary History of American Late Pleistocene Equids". Journal of Molecular Evolution. 66 (5): 533–538. Bibcode:2008JMolE..66..533O. doi:10.1007/s00239-008-9100-x. PMID 18398561. S2CID 19069554.
  50. ^ a b Cai, Dawei; Zhuowei Tang; Lu Han; et al. (2009). "Ancient DNA provides new insights into the origin of the Chinese domestic horse". Journal of Archaeological Science. 36 (3): 835–842. Bibcode:2009JArSc..36..835C. doi:10.1016/j.jas.2008.11.006.
  51. ^ a b Vilà, Carles; Jennifer A. Leonard; Anders Götherström; et al. (2001). "Widespread Origins of Domestic Horse Lineages" (PDF). Science. 291 (5503): 474–477. Bibcode:2001Sci...291..474V. doi:10.1126/science.291.5503.474. PMID 11161199. S2CID 15514589. Archived from the original (PDF) on 5 July 2020.
  52. ^ Benirschke, Poliakoff K.; N. Malouf; R. J. Low; H. Heck (16 April 1965). "Chromosome Complement: Differences between Equus caballus and Equus przewalskii". Science. 148 (3668): 382–383. doi:10.1126/science.148.3668.382. PMID 14261533. S2CID 21562513.
  53. ^ Lau, Allison; Lei Peng; Hiroki Goto; et al. (2009). "Horse Domestication and Conservation Genetics of Przewalski's Horse Inferred from Sex Chromosomal and Autosomal Sequences". Mol. Biol. Evol. 26 (1): 199–208. doi:10.1093/molbev/msn239. PMID 18931383.
  54. ^ Jansen, Thomas; Forster, Peter; Levine, Marsha A.; et al. (6 August 2002). "Mitochondrial DNA and the origins of the domestic horse". Proceedings of the National Academy of Sciences. 99 (16): 10905–10910. Bibcode:2002PNAS...9910905J. doi:10.1073/pnas.152330099. PMC 125071. PMID 12130666.
  55. ^ a b "Equus ferus ssp. przewalskii (Asian Wild Horse, Mongolian Wild Horse, Przewalski's Horse)". IUCN Red List of Threatened Species.
  56. ^ Eisenmann, Vera (1998). "Quaternary Horses: possible candidates to domestication". The Horse: its domestication, diffusion and role in past communities. Proceedings of the XIII International Congress of Prehistoric and Protohistoric Sciences, Forli, Italia, 8–14 September 1996. Vol. 1. ABACO Edizioni. pp. 27–36.
  57. ^ Pruvost, Melanie; Bellone, Rebecca; Benecke, Norbert; et al. (15 November 2011). "Genotypes of predomestic horses match phenotypes painted in Paleolithic works of cave art". Proceedings of the National Academy of Sciences. 108 (46): 18626–18630. Bibcode:2011PNAS..10818626P. doi:10.1073/pnas.1108982108. PMC 3219153. PMID 22065780.
  58. ^ Mills, Daniel S.; Nankervis, Kathryn J. (20 May 2013). Equine Behaviour: Principles and Practice. John Wiley & Sons. ISBN 978-1-118-70806-4.
  59. ^ International Commission on Zoological Nomenclature (2003). "Usage of 17 specific names based on wild species which are pre-dated by or contemporary with those based on domestic animals (Lepidoptera, Osteichthyes, Mammalia): conserved. Opinion 2027 (Case 3010)". Bull. Zool. Nomencl. 60 (1): 81–84.
  60. ^ Groves, C.; Grubb, P. (2011). Ungulate Taxonomy. Baltimore, USA: Johns Hopkins University Press. p. 179. ISBN 978-1-4214-0093-8
  61. ^ "Principles of nomenclature of zoological taxa". www.insecta.bio.spbu.ru. Archived from the original on 29 July 2017. Retrieved 28 May 2017.
  62. ^ Linnaeus, Carolus (1758). Systema naturae per regna tria naturae :secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Vol. 1 (10th ed.). Holmiae (Laurentii Salvii). p. 73. Retrieved 30 January 2009.
  63. ^ Bunzel-Drüke, Margret. "Ecological substitutes for Wild horse and Aurochs" (PDF). Natur- und Kulturlandschaft, Höxter/Jena 2001, Band 4: 10. Archived from the original (PDF) on 3 March 2016. Retrieved 28 May 2017.
  64. ^ Gentry, Anthea; Clutton-Brock, Juliet; Groves, Colin (May 2004). "The naming of wild animal species and their domestic derivates (PDF Download Available)". Journal of Archaeological Science. 31: 645–651. doi:10.1016/j.jas.2003.10.006.
  65. ^ Bunker, Emma C.; Watt, James C. Y.; Sun, Zhixin; N.Y.), Metropolitan Museum of Art (New York (2002). Nomadic Art of the Eastern Eurasian Steppes: The Eugene V. Thaw and Other New York Collections. Metropolitan Museum of Art. ISBN 978-0-300-09688-0.
  66. ^ WAZA. "Overview : WAZA: World Association of Zoos and Aquariums". www.waza.org. Archived from the original on 3 June 2017. Retrieved 28 May 2017.
  67. ^ Wallner, B.; Brem, G.; Müller, M.; Achmann, R. (2003). "Fixed nucleotide differences on the Y chromosome indicate clear divergence between Equus przewalskii and Equus caballus" (PDF). Animal Genetics. 34 (6): 453–456. doi:10.1046/j.0268-9146.2003.01044.x. PMID 14687077. S2CID 11932020. Archived from the original (PDF) on 6 July 2020.
  68. ^ Lindgren, G.; Backström, N.; Swinburne, J.; et al. (2004). "Limited number of patrilines in horse domestication". Nature Genetics. 36 (4): 335–336. doi:10.1038/ng1326. PMID 15034578.
  69. ^ Outram, A.K.; Stear, N.A.; Bendrey, R.; et al. (2009). "The earliest horse harnessing and milking". Science. 323 (5919): 1332–1335. Bibcode:2009Sci...323.1332O. doi:10.1126/science.1168594. PMID 19265018. S2CID 5126719.
  70. ^ "Rebuttal of Taylor and Barrón-Ortiz 2021 Rethinking the evidence for early horse domestication at Botai" (PDF). Retrieved 26 April 2024.
  71. ^ Outram, Alan K. (2023). "Horse domestication as a multi-centered, multi-stage process: Botai and the role of specialized Eneolithic horse pastoralism in the development of human-equine relationships". Frontiers in Environmental Archaeology. 2. doi:10.3389/fearc.2023.1134068. hdl:10871/133044.
  72. ^ Gaunitz, Charleen; Fages, Antoine; Hanghøj, Kristian; et al. (6 April 2018). "Ancient genomes revisit the ancestry of domestic and Przewalski's horses". Science. 360 (6384): 111–114. Bibcode:2018Sci...360..111G. doi:10.1126/science.aao3297. hdl:10871/31710. PMID 29472442.
  73. ^ "Ancient DNA rules out archeologists' best bet for horse domestication". ArsTechnica. 25 February 2018. Retrieved 24 June 2020.
  74. ^ Williams, Paige. "The Remarkable Comeback of Przewalski's Horse". Smithsonian.
  75. ^ "Przewalski's Horse". si.edu. Retrieved 8 June 2015.
  76. ^ Blua, Antoine (13 March 2016). "Endangered Przewalski's Horses Back On Russian Steppe". Radio Free Europe/Radio Liberty.
  77. ^ "An extraordinary return from the brink of extinction for worlds last wild horse" Archived 2006-07-22 at the Wayback Machine ZSL Living Conservation, December 19, 2005.
  78. ^ Chowdhary, Bhanu P. (22 January 2013). Equine Genomics. John Wiley & Sons. ISBN 978-1-118-52212-7.
  79. ^ The American Museum of Natural History When Is a Wild Horse Actually a Feral Horse?
  80. ^ a b "Are There Still Wild Horses?".
  81. ^ Luís, Cristina; et al. (2006). "Iberian Origins of New World Horse Breeds". Journal of Heredity. 97 (2): 107–113. doi:10.1093/jhered/esj020. PMID 16489143.
  82. ^ Rittman, Paul. "Spanish Colonial Horse and the Plains Indian Culture" (PDF). Archived from the original (PDF) on 12 April 2019. Retrieved 18 January 2015.
  83. ^ Nimmo, D. G.; Miller, K. K. (2007). "Ecological and human dimensions of management of feral horses in Australia: A review" (PDF). Wildlife Research. 34 (5): 408–417. doi:10.1071/WR06102. S2CID 26285264. Archived from the original (PDF) on 5 July 2020.
  84. ^ "Wildlife". Cumberland Island. Retrieved 20 November 2015.
  85. ^ Simons, Marlise (12 November 1995). "A Stone-Age Horse Still Roams a Tibetan Plateau". The New York Times. ISSN 0362-4331.
  86. ^ Dohner, Janet Vorwald (2001). "Equines: Natural History". In Dohner, Janet Vorwald (ed.). Historic and Endangered Livestock and Poultry Breeds. Topeka, KS: Yale University Press. pp. 400–401. ISBN 978-0-300-08880-9.
  87. ^ a b Peissel, Michel (2002). Tibet: the secret continent. Macmillan. p. 36. ISBN 978-0-312-30953-4.
  88. ^ Humi, Peter (17 November 1995). "Tibetan discovery is 'horse of a different color'". CNN. Archived from the original on 3 December 2014. Retrieved 9 September 2009.
  89. ^ McNabb, John (June 2000). "Boxgrove". Antiquity. 74 (284): 439–441. doi:10.1017/S0003598X00059548. ISSN 0003-598X.
  90. ^ Bello, Silvia M.; Parfitt, Simon A.; Stringer, Chris (September 2009). "Quantitative micromorphological analyses of cut marks produced by ancient and modern handaxes". Journal of Archaeological Science. 36 (9): 1869–1880. Bibcode:2009JArSc..36.1869B. doi:10.1016/j.jas.2009.04.014.
  91. ^ Smith, Geoff M. (October 2013). "Taphonomic resolution and hominin subsistence behaviour in the Lower Palaeolithic: differing data scales and interpretive frameworks at Boxgrove and Swanscombe (UK)". Journal of Archaeological Science. 40 (10): 3754–3767. Bibcode:2013JArSc..40.3754S. doi:10.1016/j.jas.2013.05.002.
  92. ^ Schoch, Werner H.; Bigga, Gerlinde; Böhner, Utz; et al. (December 2015). "New insights on the wooden weapons from the Paleolithic site of Schöningen". Journal of Human Evolution. 89: 214–225. Bibcode:2015JHumE..89..214S. doi:10.1016/j.jhevol.2015.08.004. PMID 26442632.
  93. ^ Urban, Brigitte; Krahn, Kim J.; Kasper, Thomas; et al. (July 2023). "Spatial interpretation of high-resolution environmental proxy data of the Middle Pleistocene Palaeolithic faunal kill site Schöningen 13 II-4, Germany". Boreas. 52 (3): 440–458. Bibcode:2023Borea..52..440U. doi:10.1111/bor.12619. ISSN 0300-9483.
  94. ^ Li, Hao; Li, Zhan-yang; Lotter, Matt G.; Kuman, Kathleen (August 2018). "Formation processes at the early Late Pleistocene archaic human site of Lingjing, China". Journal of Archaeological Science. 96: 73–84. Bibcode:2018JArSc..96...73L. doi:10.1016/j.jas.2018.05.004. hdl:2263/65160.
  95. ^ Hoffecker, John F.; Holliday, Vance T.; Stepanchuk, V.N.; Lisitsyn, S.N. (October 2018). "The hunting of horse and the problem of the Aurignacian on the central plain of Eastern Europe". Quaternary International. 492: 53–63. Bibcode:2018QuInt.492...53H. doi:10.1016/j.quaint.2018.05.044.
  96. ^ S.A. Vasil’ev. (2003). Faunal exploitation, subsistence practices and Pleistocene extinctions in Paleolithic Siberia. Deinsea, 9(1), 513–556.
  97. ^ Waters, Michael R.; Stafford, Thomas W.; Kooyman, Brian; Hills, L. V. (7 April 2015). "Late Pleistocene horse and camel hunting at the southern margin of the ice-free corridor: Reassessing the age of Wally's Beach, Canada". Proceedings of the National Academy of Sciences. 112 (14): 4263–4267. Bibcode:2015PNAS..112.4263W. doi:10.1073/pnas.1420650112. ISSN 0027-8424. PMC 4394292. PMID 25831543.
  98. ^ Librado, Pablo; Khan, Naveed; Fages, Antoine; et al. (28 October 2021). "The origins and spread of domestic horses from the Western Eurasian steppes". Nature. 598 (7882): 634–640. Bibcode:2021Natur.598..634L. doi:10.1038/s41586-021-04018-9. ISSN 0028-0836. PMC 8550961. PMID 34671162.

Bibliography

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